Draft Key of the Doryctinae genera of the world for each geographic regions

This key is only a draft version provided to test the site. It has not been tested enough to confirm its quality. Moreover some errors could be present. So, if you wish to use it, please take in mind that all your identifications need to be seen with cautions and verified with additional sources!


The following genera have not been included because I lack data : Cecidospathius Kieffer & Jörgensen,1910; Celereon Say,1936; Doryctomorpha Asmead, 1900; Esterella Pagliano & Scaramozzino,1990; Metaspathius Brues, 1922; Sinaodoryctes Chen & Shi, 2004 (Key version 08/10/2017).


1(0).
  • Nearctic ... 2
  • Eastern Palearctic... 31
  • Western Palaearctic... 62
  • Ethiopian... 87
  • Neotropical... 133
  • Oriental... 247
  • Australasian... 297
  • Oceania... 353

2(1).
  • Scape (sub)equal to first flagellomere ... 3
  • Scape shorter than first flagellomere... 5

3(2).
  • Apical lobe of scape absent; occipital carina present dorsally and laterally; apical margin of scape and pedicel without dense cluster of setae; third labial palp segment as long as the second ... 4
  • Apical lobe of scape present (apically or sub-apically); occipital carina absent; apical margin of scape and pedicel with dense cluster of setae; third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... Liobracon Szépligeti, 1901

4(3).
  • Occipital carina meeting hypostomal carina; notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86); scutellum flat; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)) ... Stenocorse Marsh, 1968
  • Occipital carina not meeting hypostomal carina; notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14); scutellum convex (e.g. swollen, conical, sharply raised); metapleuron glabrous or with a very sparse/rare discret pubescence... Rhoptrocentrus Marshall, 1897

5(2).
  • Maxillary palpi six-segmented; labial palpi four-segmented ... 6
  • Maxillary palpi five-segmented; labial palpi three-segmented... 29

6(5).
  • Petiole without median carinae ... 7
  • Petiole with median carinae... 9

7(6).
  • Scutellum flat; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B); spiral stucture of venom reservoir normal ... 8
  • Scutellum convex (e.g. swollen, conical, sharply raised); propodeum clearly convex; fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F); spiral stucture of venom reservoir posteriorly much courser than anteriorly... Aptenobracon Marsh, 1965

8(7).
  • Pronotum with convex lobe; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; propodeal carinated areas present at least basally (e.g. Fig. 2E); hind femora not swollen (at least 3x longer than wide) ... Curtisella Spinola, 1853
  • Pronotum dorsally without modifications; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; propodeal carinated areas completely absent; hind femora swollen, at most 2.6x longer than wide... Pambolidea Ashmead, 1900

9(6).
  • Third labial palp segment as long as the second; hind coxa dorsally without teeth (e.g. Fig. 5A, B, E); raised ridge between antennae absent; ctenidia minor type only ... 10
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113); hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D); raised ridge between antennae present; ctenidia major type present... Odontobracon Cameron, 1887

10(9).
  • Sixth tergite visible; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B) ... 11
  • Sixth tergite hidden (by the fifth tergite); fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F)... Rhaconotus Ruthe, 1854

11(10).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14) ... 12
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86)... 28

12(11).
  • Fore femur with more or less distinct dorsal blister ... 13
  • Fore femur without dorsal blister... 16

13(12).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence; first flagellar segment straigth; frons not excavated (flat or convex); mid tibia without spines on antero dorsal side ... 14
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); first flagellar segment more or less curved; frons excavated (concave, sometimes weakly); mid tibia with spines on antero dorsal side... Doryctinus Roman, 1910

14(13).
  • Hind femora with more or less distinct dorsal blister; comprising two parts ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts... 15

15(14).
  • Pronotum carina(e) or keel absent or indistinct; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; mid femora without dorsal blister ... Parallorhogas Marsh, 1993
  • Pronotum carina(e) or keel present (laterally and-or dorsally); mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); mid femora with more or less distinct dorsal blister... Ontsira Cameron, 1900

16(12).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... 17
  • Acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E)... 26
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Spathius Nees, 1918

17(16).
  • Basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64) ... 18
  • Basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63)... 20

18(17).
  • Hind femora swollen, at most 2.6x longer than wide; frons not excavated (flat or convex); frons without lateral protuberances; subctenidial setae simple ... 19
  • Hind femora not swollen (at least 3x longer than wide); frons excavated (concave, sometimes weakly); frons with lateral protuberances; subctenidial setae bifurcate... Dendrosoter Wesmael, 1838

19(18).
  • Vertex not striate; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); hind tibia with (comb or area) of dense setae at inner apex; comprising three parts ... Hemidoryctes Belokobylskij, 1993
  • Vertex striate; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); hind tibia without (comb or area) of dense setae at inner apex; comprising two parts... Pambolidea Ashmead, 1900

20(17).
  • Hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C) ... 21
  • Hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K)... 23

21(20).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); position of the valvillar insertion medial; subctenidial setae not or hardly flattened; single, large crescentic bar-like structure just distal to valvillus present ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); position of the valvillar insertion close to the dorsal edge of the egg canal; subctenidial setae distinctly flattened; single, large crescentic bar-like structure just distal to valvillus absent... 22

22(21).
  • Occipital carina meeting hypostomal carina; vertex not striate; pronotum dorsally without modifications; pronotum carina(e) or keel present (laterally and-or dorsally) ... Allorhogas Gahan, 1912
  • Occipital carina not meeting hypostomal carina; vertex striate; pronotum with convex lobe; pronotum carina(e) or keel absent or indistinct... Monolexis Foerster, 1862

23(20).
  • Pronotum short (nearly invisible in dorsal view); pronotum dorsally without modifications; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)) ... Glyptocolastes Ashmead, 1900
  • Pronotum normal to long (visible in dorsal view); pronotum with convex lobe; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; metapleuron glabrous or with a very sparse/rare discret pubescence... 24

24(23).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); hind tibia without (comb or area) of dense setae at inner apex; second tergite with apical lenticulate area (e.g. Fig. 6F, I) (often formed by the meeting of second suture and groove on third tergite when present) ... 25
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; propodeum clearly convex; hind tibia with (comb or area) of dense setae at inner apex; second tergite without apical lenticulate area (e.g. Fig. 6B, E)... Leluthia Cameron, 1887 (female)

25(24).
  • Scutellum without lateral carinae; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); first flagellar segment straigth; third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area) ... Leluthia Cameron, 1887 (male)
  • Scutellum with high subvertical lateral lobes/carinae; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); first flagellar segment more or less curved; third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G)... Callihormius Ashmead, 1900b

26(16).
  • Vertex not striate; vein 2-1A of fore wing not shortened; petiole without basolateral process (pointed wing-like, spine-like projections) near base ... 27
  • Vertex striate; vein 2-1A of fore wing more or less shortened; petiole with basolateral process (pointed wing-like, spine-like projections) near base... Ptesimogaster Marsh, 1965

27(26).
  • Pronotum with convex lobe; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; first flagellar segment longer than second (e.g. Fig. 1H, I); second radiomedial vein (r-m) of fore wing with wide bulla (e.g. Fig. 3d) ... Callihormius Ashmead, 1900b
  • Pronotum dorsally without modifications; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; first flagellar segment (sub)equal to second (e.g. Fig. 1L); second radiomedial vein (r-m) of fore wing largely tubular (e.g. Fig. 4H; Marsh, 1997: figs 16, 18, 30)... Pioscelus Muesebeck & Walkley, 1951

28(11).
  • Malar suture absent or indistinct; vertex not striate; pronotum with convex lobe; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum ... Callihormius Ashmead, 1900b
  • Malar suture present; vertex striate; pronotum dorsally without modifications; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum... Percnobraconoides Marsh, 1989

29(5).
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); metanotum with a short and acute erected spine medially; propodeal bridge between abdominal and coxal foramina absent; second abscissa of costal vein in hind wing usually present (sclerotized) ... 30
  • First flagellar segment shorter than second; metanotum without any protuberances or spines; propodeal bridge between abdominal and coxal foramina present; second abscissa of costal vein in hind wing absent... Aivalykus Nixon, 1938

30(29).
  • Wings present; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Ecphylus Foerster, 1862 (winged forms)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

31(1).
  • Scape (sub)equal to first flagellomere ... 32
  • Scape shorter than first flagellomere... 36

32(31).
  • Petiole median carinae very short (less than or equal to the third of petiole length) ... Zombrus Marshall, 1897
  • Petiole median carinae reaching, or nearly, the half length of the petiole... 33
  • Petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... 34

33(32).
  • Hind femora swollen, at most 2.6x longer than wide; eyes more or less emarginated on their inner sides; vein r of fore wing arising distinctly before middle of pterostigma; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64) ... Cryptontsira Belokobylskij, 2008
  • Hind femora not swollen (at least 3x longer than wide); eyes not emarginated; vein r of fore wing arising distinctly after(behind) middle of pterostigma; basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63)... Spathiomorpha Tobias, 1976

34(32).
  • Notauli not meeting each other ... Nipponecphylus Belokobylskij & Konishi, 2001 (males)
  • Notauli meeting before the scutelar sulcus... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Notauli meeting a sculptured area before scutelar sulcus... 35

35(34).
  • Scape less than two times longer than maximally wide; metanotum without any protuberances or spines; vein r of fore wing arising submedially from pterostigma; second radiomedial vein (r-m) of fore wing present ... Rhoptrocentrus Marshall, 1897
  • Scape two or more times longer than maximally wide; metanotum with a short (obtuse) median tubercle; vein r of fore wing arising distinctly before middle of pterostigma; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated... Hecabolus Curtis, 1837

36(31).
  • Apical lobe of scape absent ... 37
  • Apical lobe of scape present (apically or sub-apically)... Asiaheterospilus Belokobylskij & Konishi, 2001

37(36).
  • Maxillary palpi six-segmented; labial palpi four-segmented ... 38
  • Maxillary palpi five-segmented; labial palpi three-segmented... 60

38(37).
  • Sixth tergite visible; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B) ... 39
  • Sixth tergite hidden (by the fifth tergite); fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F)... Rhaconotus Ruthe, 1854

39(38).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence ... 40
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body))... 57

40(39).
  • Dorsope of first metasomal tergite present and more or less distinct ... 41
  • Dorsope of first metasomal tergite absent or indistinct... 56

41(40).
  • Mesoscutum surface of median and lateral lobes smooth and shining ... 42
  • Mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa)... Heterospilus Halliday, 1836
  • Mesoscutum surface of median and lateral lobes more or less sculptured... 44

42(41).
  • Nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F); recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); scape longer dorsally than ventrally; mid femora without dorsal blister ... 43
  • Nervellus (cu-a) of fore wing interstitial; recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); scape as long as ventrally than dorsally; mid femora with more or less distinct dorsal blister... Hypodoryctes Kokujev, 1900

43(42).
  • Pronotum short (nearly invisible in dorsal view); pronotum carina(e) or keel absent or indistinct; brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); vein 2-1A of fore wing more or less shortened ... Parallorhogas Marsh, 1993
  • Pronotum normal to long (visible in dorsal view); pronotum carina(e) or keel present (laterally and-or dorsally); brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); vein 2-1A of fore wing not shortened... Doryctes (Doryctes + Neodoryctes) Haliday, 1836

44(41).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... 45
  • Fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B)... Leluthia Cameron, 1887 (male)

45(44).
  • Basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F) ... 46
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J)... 53

46(45).
  • Fore femur with more or less distinct dorsal blister; mid femora with more or less distinct dorsal blister ... 47
  • Fore femur without dorsal blister; mid femora without dorsal blister... 49

47(46).
  • Mid tibia without spines on antero dorsal side; male genitalia with basal lobe of basal ring (e.g. Fig. 7B, G) ... 48
  • Mid tibia with spines on antero dorsal side; male genitalia without basal lobe of basal ring (e.g. Fig. 8A, C)... Rhacontsira Belokobylskij, 1998

48(47).
  • Hind femora with more or less distinct dorsal blister; comprising two parts; bars anterior to valvillus or valvillar zone absent; bar posterior to valvillus or valvillar zone absent ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts; bars anterior to valvillus or valvillar zone present; bar posterior to valvillus or valvillar zone present... Ontsira Cameron, 1900

49(46).
  • Ovipositor nodus without visible node (but lower valves serrate) ... Caenophanes Foerster, 1862
  • Ovipositor nodus double, with a second node weakly developed (e.g. Quicke et al., 1992a: fig. 12)... Dendrosotinus Telenga, 1941
  • Ovipositor nodus double (or more as Cryptodoryctes), with both nodus well developed (e.g. Quicke et al., 1992a: figs 1-8)... 50

50(49).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64) ... 51
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63)... 52

51(50).
  • Pronotum with convex lobe; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent) ... Pareucorystes Tobias, 1961
  • Pronotum dorsally without modifications; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present)... Dendrosoter Wesmael, 1838

52(50).
  • Petiole median carinae very short (less than or equal to the third of petiole length) ... Monolexis Foerster, 1862
  • Petiole median carinae reaching, or nearly, the half length of the petiole... Hecabalodes Wilkinson, 1929
  • Petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... Leluthia Cameron, 1887 (female)

53(45).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron) ... 54
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron)... 55

54(53).
  • First flagellar segment surface dorso-ventrally not compressed; scutellum flat; first flagellar segment straigth; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated) ... Leluthia (Euhecabolodes) Tobias, 1962
  • First flagellar segment surface dorso-ventrally compressed (sometimes strongly); scutellum convex (e.g. swollen, conical, sharply raised); first flagellar segment more or less curved; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully)... Sonanus Belokobylskij & Konishi, 2001

55(53).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Eodendrus Belokobylskij, 1998
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Hecabalodes Wilkinson, 1929
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein)... Polystenus Förster, 1862

56(40).
  • Vertex not striate; pronotum with convex lobe; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Platyspathius Viereck, 1911
  • Vertex striate; pronotum dorsally without modifications; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Spathius (Stenophasmus) Smith, 1858

57(39).
  • Dorsal tentorial pits near the antennal sockets absent ... 58
  • Dorsal tentorial pits near the antennal sockets present between antenal sockets and eyes... Neurocrassus Snoflak, 1945

58(57).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); propodeal carinated areas present at least basally (e.g. Fig. 2E); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 59
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); propodeal carinated areas completely absent; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Polystenus Förster, 1862

59(58).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F); fore femur without dorsal blister ... Spathius Nees, 1918
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F); basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J); fore femur with more or less distinct dorsal blister... Hecabolomorpha Belokobylskij & Chen, 2006

60(37).
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); metanotum with a short and acute erected spine medially; propodeal bridge between abdominal and coxal foramina absent; second abscissa of costal vein in hind wing usually present (sclerotized) ... 61
  • First flagellar segment shorter than second; metanotum without any protuberances or spines; propodeal bridge between abdominal and coxal foramina present; second abscissa of costal vein in hind wing absent... Aivalykus Nixon, 1938

61(60).
  • Wings present; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Ecphylus Foerster, 1862 (winged forms)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

62(1).
  • Metasoma of male not depressed, rather long, not specially curved ... 63
  • Metasoma of male not depressed dorsoventrally, relatively short, curved down apically... 86
  • Metasoma of male depressed dorsoventrally, distinctly longer than head and mesosoma combined, not curved down apically... Sycosoter Picard & Lichtenstein, 1917

63(62).
  • Third labial palp segment as long as the second ... 64
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... 85

64(63).
  • Scape (sub)equal to first flagellomere ... 65
  • Scape shorter than first flagellomere... 67

65(64).
  • Metanotum without any protuberances or spines ... 66
  • Metanotum with a short and acute erected spine medially... Spathiomorpha Tobias, 1976
  • Metanotum with a short (obtuse) median tubercle... Hecabolus Curtis, 1837

66(65).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; scutellum convex (e.g. swollen, conical, sharply raised); second radiomedial vein (r-m) of fore wing present; fore wing hyaline ... Rhoptrocentrus Marshall, 1897
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; scutellum flat; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; fore wing banded... Pseudorhaconotus van Achterberg & Shaw, 2010

67(64).
  • Hind femur without teeth on ventral side; hind coxa dorsally without teeth (e.g. Fig. 5A, B, E) ... 68
  • Hind femur with more than one tooth (often one or two and several smaller) on ventral side; hind coxa with one to several teeth ventrally... Euscelinus Westwood, 1882

68(67).
  • Sixth tergite visible; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B) ... 69
  • Sixth tergite hidden (by the fifth tergite); fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F)... Rhaconotus Ruthe, 1854

69(68).
  • Fourth-sixth metasomal tergites of males simple, submarginal lateral carinae absent ... 70
  • Fourth-sixth metasomal tergites of males submarginal lateral carinae present (e.g. Fig. 7E)... 83

70(69).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence; subctenidial setae simple ... 71
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); subctenidial setae trifurcate or more divided... Spathius Nees, 1918

71(70).
  • Dorsope of first metasomal tergite present and more or less distinct ... 72
  • Dorsope of first metasomal tergite absent or indistinct... Platyspathius Viereck, 1911

72(71).
  • Fore femur with more or less distinct dorsal blister ... 73
  • Fore femur without dorsal blister... 76

73(72).
  • Hind femora with more or less distinct dorsal blister; comprising two parts ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts... 74

74(73).
  • Mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; parameres wide and roundly triangular; bars anterior to valvillus or valvillar zone absent ... 75
  • Mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); parameres considerably narrowed along whole length (e.g. Fig. 8E); bars anterior to valvillus or valvillar zone present... Ontsira Cameron, 1900

75(74).
  • Occipital carina meeting hypostomal carina; recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); nervellus (cu-a) of fore wing interstitial; brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present) ... Hypodoryctes Kokujev, 1900
  • Occipital carina not meeting hypostomal carina; recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F); brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent)... Parallorhogas Marsh, 1993

76(72).
  • Ovipositor nodus without visible node (but lower valves serrate) ... Caenophanes Foerster, 1862
  • Ovipositor nodus double, with a second node weakly developed (e.g. Quicke et al., 1992a: fig. 12)... Dendrosotinus Telenga, 1941
  • Ovipositor nodus double (or more as Cryptodoryctes), with both nodus well developed (e.g. Quicke et al., 1992a: figs 1-8)... 77

77(76).
  • Hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C) ... 78
  • Hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K)... 80

78(77).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); position of the valvillar insertion medial; subctenidial setae not or hardly flattened; single, large crescentic bar-like structure just distal to valvillus present ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); position of the valvillar insertion close to the dorsal edge of the egg canal; subctenidial setae distinctly flattened; single, large crescentic bar-like structure just distal to valvillus absent... 79

79(78).
  • Occipital carina meeting hypostomal carina; vertex not striate; pronotum dorsally without modifications; pronotum carina(e) or keel present (laterally and-or dorsally) ... Allorhogas Gahan, 1912
  • Occipital carina not meeting hypostomal carina; vertex striate; pronotum with convex lobe; pronotum carina(e) or keel absent or indistinct... Monolexis Foerster, 1862

80(77).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64); comprising two parts ... Pareucorystes Tobias, 1961
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63); comprising three parts... 81

81(80).
  • Pronotum with convex lobe; hind femora not swollen (at least 3x longer than wide); petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole); male genitalia without basal lobe of basal ring (e.g. Fig. 8A, C) ... 82
  • Pronotum dorsally without modifications; hind femora swollen, at most 2.6x longer than wide; petiole median carinae reaching, or nearly, the half length of the petiole; male genitalia with basal lobe of basal ring (e.g. Fig. 7B, G)... Hecabalodes Wilkinson, 1929

82(81).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B); propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... Leluthia Cameron, 1887 (male)
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F); fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly; propodeum clearly convex... Leluthia Cameron, 1887 (female)

83(69).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Dendrosoter Wesmael, 1838
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... 84
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein)... Polystenus Förster, 1862

84(83).
  • First flagellar segment longer than second (e.g. Fig. 1H, I); ocelar triangle with base larger than its sides; pronotum normal to long (visible in dorsal view); pronotum with convex lobe ... Leluthia Cameron, 1887 (female)
  • First flagellar segment shorter than second; ocelar triangle with base shorter than its sides; pronotum short (nearly invisible in dorsal view); pronotum dorsally without modifications... Caenopachys Förster, 1862 female

85(63).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; pronotum dorsally without modifications; vertex striate; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated ... Mimodoryctes Belokobylskij, 2001
  • Recurrent vein (m-cu) of hind wing strongly curved to apex of wing (fully sclerified usually); pronotum with convex lobe; vertex not striate; second radiomedial vein (r-m) of fore wing present... Zombrus Marshall, 1897

86(62).
  • Wings present; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Ecphylus Foerster, 1862 (winged forms)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

87(1).
  • Propodeal bridge between abdominal and coxal foramina absent ... 88
  • Propodeal bridge between abdominal and coxal foramina present... 131

88(87).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14) ... 89
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86)... 130

89(88).
  • Sixth tergite visible ... 90
  • Sixth tergite hidden (by the fifth tergite)... 129

90(89).
  • Pronope present ... Cryptodoryctes Belokobylskij & Quicke, 2000
  • Pronope absent... 91

91(90).
  • Prescutellar depression (scutellar sulcus) long and narrow or of medium length ... 92
  • Prescutellar depression (scutellar sulcus) considerably shortened (e.g. Achterberg, 1995: figs 35, 55)... Hemispathius Belokobylskij & Quicke, 2000

92(91).
  • Third labial palp segment as long as the second ... 93
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... 128
  • Third labial palp segment longer than the second... genA

93(92).
  • Fore tibial spines absent ... Leptorhaconotus Granger, 1949
  • Fore tibial spines present (e.g. Fig. 5G; Marsh, 1997: figs 65-67)... 94

94(93).
  • Apical lobe of scape absent; scape less than two times longer than maximally wide ... 95
  • Apical lobe of scape present (apically or sub-apically); scape two or more times longer than maximally wide... Rinamba Cameron,1912

95(94).
  • Maxillary palpi six-segmented; labial palpi four-segmented; second abscissa of costal vein in hind wing absent; metasoma of male not depressed, rather long, not specially curved ... 96
  • Maxillary palpi five-segmented; labial palpi three-segmented; second abscissa of costal vein in hind wing usually present (sclerotized); metasoma of male not depressed dorsoventrally, relatively short, curved down apically... 127

96(95).
  • First flagellar segment surface dorso-ventrally not compressed ... 97
  • First flagellar segment surface dorso-ventrally compressed (sometimes strongly)... 125

97(96).
  • Dorsope of first metasomal tergite present and more or less distinct ... 98
  • Dorsope of first metasomal tergite absent or indistinct... 122

98(97).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... 99
  • Fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B)... 118
  • Fourth-sixth metasomal tergites of males strongly sclerotised, distinctly convex, entirely sculptured, sixth tergite enlarged, covered following tergites... Bathycentor Saussure, 1892

99(98).
  • Fourth-sixth metasomal tergites of males simple, submarginal lateral carinae absent ... 100
  • Fourth-sixth metasomal tergites of males submarginal lateral carinae present (e.g. Fig. 7E)... 117

100(99).
  • Hind coxa dorsally without teeth (e.g. Fig. 5A, B, E) ... 101
  • Hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D)... Priosphys Enderlein, 1920

101(100).
  • Mid tibia without spines on antero dorsal side ... 102
  • Mid tibia with spines on antero dorsal side... 114

102(101).
  • Fore femur with more or less distinct dorsal blister ... 103
  • Fore femur without dorsal blister... 105

103(102).
  • Hind femora with more or less distinct dorsal blister; comprising two parts ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts... 104

104(103).
  • Pronotum carina(e) or keel absent or indistinct; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; mid femora without dorsal blister ... Parallorhogas Marsh, 1993
  • Pronotum carina(e) or keel present (laterally and-or dorsally); mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); mid femora with more or less distinct dorsal blister... Ontsira Cameron, 1900

105(102).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... 106
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Spathius Nees, 1918

106(105).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 107
  • Propodeal carinated areas completely absent... 111

107(106).
  • Second tergite without apical lenticulate area (e.g. Fig. 6B, E) ... 108
  • Second tergite with apical lenticulate area (e.g. Fig. 6F, I) (often formed by the meeting of second suture and groove on third tergite when present)... Bathycentor Saussure, 1892

108(107).
  • Head with sparse pilosity (short or long, sometimes in punctuations) ... 109
  • Head with long pilosity somewhere (face, vertex, fully,...)... Dendrosotinus Telenga, 1941

109(108).
  • First radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; fore wing of male without sclerotized enlargement, veins (3-SR, SR1, 2-M and 3-M) slender; basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 110
  • First radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; fore wing of male with sclerotized enlargement on some veins (as 3-SR, SR1, 2-M and 3-M); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Bulbonervus Shenefelt, 1969

110(109).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); frons not excavated (flat or convex); position of the valvillar insertion medial; bars anterior to valvillus or valvillar zone present ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); frons excavated (concave, sometimes weakly); position of the valvillar insertion close to the dorsal edge of the egg canal; bars anterior to valvillus or valvillar zone absent... Allorhogas Gahan, 1912

111(106).
  • Vertex not striate ... 112
  • Vertex striate... 113

112(111).
  • Mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); second radiomedial vein (r-m) of fore wing present; recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally) ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS)... Hecabalodes Wilkinson, 1929

113(111).
  • Vein 2-1A of fore wing not shortened; hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64); hind tibia without (comb or area) of dense setae at inner apex ... Terate Nixon, 1943
  • Vein 2-1A of fore wing more or less shortened; hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63); hind tibia with (comb or area) of dense setae at inner apex... Monolexis Foerster, 1862

114(101).
  • First flagellar segment longer than second (e.g. Fig. 1H, I) ... 115
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L)... Eodendrus Belokobylskij, 1998
  • First flagellar segment shorter than second... 116

115(114).
  • Occipital carina meeting hypostomal carina; mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); metanotum with a short (obtuse) median tubercle ... Doryctinus Roman, 1910
  • Occipital carina not meeting hypostomal carina; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; metanotum without any protuberances or spines... Doryctoproctus Belokobylskij, 2004

116(114).
  • Vertex not striate; acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E); ovipositor sheath with normal, short and thin setae ... genB male
  • Vertex striate; acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); ovipositor sheath sparsely covered by long stout setae... genB female

117(99).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Dendrosoter Wesmael, 1838
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Caenopachys Förster, 1862 female
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein)... Bathycentor Saussure, 1892

118(98).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Grangerdoryctes Belokobylskij, 2004
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Ipodoryctes Granger, 1949
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein)... 119

119(118).
  • Vertex not striate; hind coxa dorsally without teeth (e.g. Fig. 5A, B, E); petiole without basolateral process (pointed wing-like, spine-like projections) near base ... 120
  • Vertex striate; hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D); petiole with basolateral process (pointed wing-like, spine-like projections) near base... Odontodoryctes Granger, 1949

120(119).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E); fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67); first flagellar segment longer than second (e.g. Fig. 1H, I); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 121
  • Propodeal carinated areas completely absent; fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G); first flagellar segment shorter than second; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Gymnobracon Szépligeti, 1902

121(120).
  • Pronotum with convex lobe; scutellum convex (e.g. swollen, conical, sharply raised); first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent) ... Neoheterospilus (Neoheterospilus) Belokobylskij, 2006
  • Pronotum dorsally without modifications; scutellum flat; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present)... Bathycentor Saussure, 1892

122(97).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; total length of maxillary palps reaching mid coxae; sternaulus (precoxal sulcus) absent; mid tibia with spines on antero dorsal side ... genC male
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; total length of maxillary palps not reaching mid coxae; sternaulus (precoxal sulcus) present; mid tibia without spines on antero dorsal side... 123

123(122).
  • Pronotum short (nearly invisible in dorsal view); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); clypeus without lower flange; propodeum clearly convex ... Spathioplites Fisher, 1962
  • Pronotum normal to long (visible in dorsal view); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); clypeus with lower flange more or less developped; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... 124

124(123).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence; acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E); fore femur with more or less distinct dorsal blister; mid femora with more or less distinct dorsal blister ... Platyspathius Viereck, 1911
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); fore femur without dorsal blister; mid femora without dorsal blister... Spathius Nees, 1918

125(96).
  • Mesoscutum surface of median and lateral lobes smooth and shining; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B) ... Paradoryctes Granger, 1949
  • Mesoscutum surface of median and lateral lobes more or less sculptured; metapleuron glabrous or with a very sparse/rare discret pubescence; basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F); fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly... 126

126(125).
  • Pronotum short (nearly invisible in dorsal view); pronotum dorsally without modifications; pronotum carina(e) or keel present (laterally and-or dorsally); scape shorter than first flagellomere ... Dendrosotinus Telenga, 1941
  • Pronotum very long (in dorsal view); pronotum with convex lobe; pronotum carina(e) or keel absent or indistinct; scape (sub)equal to first flagellomere... Doryctophasmus Enderlein, 1912

127(95).
  • Wings present; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Ecphylus Foerster, 1862 (winged forms)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

128(92).
  • Scape (sub)equal to first flagellomere; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron glabrous or with a very sparse/rare discret pubescence; propodeal carinated areas completely absent ... Zombrus Marshall, 1897
  • Scape shorter than first flagellomere; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); propodeal carinated areas present at least basally (e.g. Fig. 2E)... Ivondrovia Shenefelt & Marsh, 1976

129(89).
  • Scape (sub)equal to first flagellomere; occipital carina meeting hypostomal carina; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); propodeum with lateral tubercles (as high as large at base) ... Rhaconotus (Pararhacon) Belokobylskij, 2004
  • Scape shorter than first flagellomere; occipital carina not meeting hypostomal carina; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); propodeum without lateral tubercles... Rhaconotus Ruthe, 1854

130(88).
  • First flagellar segment longer than second (e.g. Fig. 1H, I); vertex striate; mesoscutum overhanging over the pronotum; metanotum with a short (obtuse) median tubercle ... Afrospathius Belokobylskij & Quicke, 2000
  • First flagellar segment shorter than second; vertex not striate; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; metanotum without any protuberances or spines... Toka Nixon, 1943

131(87).
  • First flagellar segment longer than second (e.g. Fig. 1H, I) ... Pseudodoryctes Szépligeti, 1915
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L)... 132
  • First flagellar segment shorter than second... Aivalykus Nixon, 1938

132(131).
  • Pronope present; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; first flagellar segment surface dorso-ventrally not compressed; ocelar triangle with base larger than its sides ... Stephanospathius Belokobylskij, 1993
  • Pronope absent; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; first flagellar segment surface dorso-ventrally compressed (sometimes strongly); ocelar triangle with base shorter than its sides... Schlettereriella Szépligeti, 1904

133(1).
  • Pronope present ... 134
  • Pronope absent... 135

134(133).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; third labial palp segment as long as the second; petiole median carinae reaching, or nearly, the half length of the petiole; malar suture present ... Mexiare Belokobylskij, Samaca-Saenz & Zaldivar-Riveron, 2015
  • Recurrent vein (m-cu) of hind wing strongly curved to apex of wing (fully sclerified usually); third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113); petiole median carinae very short (less than or equal to the third of petiole length); malar suture absent or indistinct... Nervellius Roman, 1924

135(133).
  • Metasoma of male not depressed, rather long, not specially curved; second abscissa of costal vein in hind wing absent ... 136
  • Metasoma of male not depressed dorsoventrally, relatively short, curved down apically; second abscissa of costal vein in hind wing usually present (sclerotized)... 246

136(135).
  • Propodeum without median process ... 137
  • Propodeum with a median raised tooth or tubercle... Coiba Marsh, 1993
  • Propodeum with a medio-longitudinal raised lamella... Bohartiellus Marsh, 1983

137(136).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 138
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 220

138(137).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14) ... 139
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86)... 215

139(138).
  • Fore femur with more or less distinct dorsal blister ... 140
  • Fore femur without dorsal blister... 154

140(139).
  • Petiole without median carinae ... 141
  • Petiole with median carinae... 142

141(140).
  • Scape (sub)equal to first flagellomere; first flagellar segment longer than second (e.g. Fig. 1H, I); maxillary palpi six-segmented; third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113) ... Monarea Szépligeti, 1904
  • Scape shorter than first flagellomere; first flagellar segment (sub)equal to second (e.g. Fig. 1L); maxillary palpi five-segmented; third labial palp segment as long as the second... Sergey Martínez, Lázaro, Pedraza-Lara & Zaldívar-Riverón, 2016

142(140).
  • Scape less than two times longer than maximally wide; second metasomal suture present (e.g. Fig. 7C, D, F) ... 143
  • Scape two or more times longer than maximally wide; second metasomal suture indistinct, to largely or entirely absent (e.g. Figs 6B, E, 7E)... Venifurca Gadelha, Nunes & Zaldívar-Riverón, 2016

143(142).
  • Maxillary palpi six-segmented; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B) ... 144
  • Maxillary palpi five-segmented; fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F)... Ondigus Braet, Barbalhoa & van Achterberg, 2003

144(143).
  • Propodeal bridge between abdominal and coxal foramina absent ... 145
  • Propodeal bridge between abdominal and coxal foramina present... Neostaphius Braet, Barbalhoa & van Achterberg, 2003

145(144).
  • Dorsope of first metasomal tergite present and more or less distinct ... 146
  • Dorsope of first metasomal tergite absent or indistinct... Concurtisella Roman, 1924

146(145).
  • Scape as long as ventrally than dorsally ... 147
  • Scape longer dorsally than ventrally... 151

147(146).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base ... 148
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified)... 150

148(147).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D); fore wing more or less infuscate (sometimes with pale spots inside); petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area) ... Barbalhoa Marsh, 2002
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C); fore wing hyaline; petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G)... 149

149(148).
  • Vertex not striate; mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Hypodoryctes Kokujev, 1900
  • Vertex striate; mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Coiba Marsh, 1993

150(147).
  • Vertex not striate; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); vein r of fore wing arising submedially from pterostigma; fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67) ... Semirhytus Szépligeti, 1902
  • Vertex striate; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); vein r of fore wing arising distinctly before middle of pterostigma; fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G)... Johnsonius Marsh, 1993

151(146).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly; valvillus of lower ovipositor valve present and well developped ... 152
  • Fourth-sixth metasomal tergites of males strongly sclerotised, distinctly convex, entirely sculptured, sixth tergite enlarged, covered following tergites; valvillus of lower ovipositor valve absent... Pedinotus Szépligeti, 1902

152(151).
  • Metanotum without any protuberances or spines; metapleuron glabrous or with a very sparse/rare discret pubescence; first flagellar segment straigth; frons not excavated (flat or convex) ... 153
  • Metanotum with a short (obtuse) median tubercle; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); first flagellar segment more or less curved; frons excavated (concave, sometimes weakly)... Doryctinus Roman, 1910

153(152).
  • Occipital carina meeting hypostomal carina; petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole); mid femora with more or less distinct dorsal blister; hind femora with more or less distinct dorsal blister ... Heterospilus Halliday, 1836
  • Occipital carina not meeting hypostomal carina; petiole median carinae very short (less than or equal to the third of petiole length); mid femora without dorsal blister; hind femora without dorsal blister... Parallorhogas Marsh, 1993

154(139).
  • Basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F) ... 155
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J)... 205

155(154).
  • Scutellum flat ... 156
  • Scutellum convex (e.g. swollen, conical, sharply raised)... 200

156(155).
  • Propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 157
  • Propodeum clearly convex... 193

157(156).
  • Hind tibia without (comb or area) of dense setae at inner apex ... 158
  • Hind tibia with (comb or area) of dense setae at inner apex... 182

158(157).
  • Basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64) ... 159
  • Basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63)... 168

159(158).
  • Hind wing with the nervellus (cu-a) vein inclivous (e.g. Figs 3E, 4B)(inclinated to base of wing) ... 160
  • Hind wing without the nervellus (cu-a) vein (e.g. Fig. 4A, E)... 165

160(159).
  • Mesoscutum surface of median and lateral lobes smooth and shining ... 161
  • Mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa)... Tuberatra Gadelha, Nunes & De Oliveira, 2016
  • Mesoscutum surface of median and lateral lobes more or less sculptured... 164

161(160).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base ... 162
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified)... Glaucia Braga, Barbalho & Penteado-Dias, 2002
  • Recurrent vein (m-cu) of hind wing strongly curved to apex of wing (fully sclerified usually)... Nervellius philippus Braet, 2014

162(161).
  • Petiole median carinae very short (less than or equal to the third of petiole length) ... 163
  • Petiole median carinae reaching, or nearly, the half length of the petiole... Megaloproctus Schulz, 1906
  • Petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... Histeromeroides Marsh, 1993

163(162).
  • Vertex not striate; propodeal carinated areas present at least basally (e.g. Fig. 2E); second radiomedial vein (r-m) of fore wing with wide bulla (e.g. Fig. 3d); fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G) ... Rinamba Cameron,1912
  • Vertex striate; propodeal carinated areas completely absent; second radiomedial vein (r-m) of fore wing largely tubular (e.g. Fig. 4H; Marsh, 1997: figs 16, 18, 30); fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67)... Pannuceus Marsh, 2002

164(160).
  • Vertex not striate; scape two or more times longer than maximally wide; scape (sub)equal to first flagellomere; propodeal carinated areas completely absent ... Verae Marsh, 1993
  • Vertex striate; scape less than two times longer than maximally wide; scape shorter than first flagellomere; propodeal carinated areas present at least basally (e.g. Fig. 2E)... Bolivar Zaldívar-Riverón & Rodríguez-Jiménez, 2013

165(159).
  • First radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; radial (marginal) cell of fore wing distally closed; hind wing without the recurrent vein (m-cu); sternaulus (precoxal sulcus) present ... 166
  • First radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; radial (marginal) cell of fore wing open; hind wing with the recurrent vein (m-cu); sternaulus (precoxal sulcus) absent... 167

166(165).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E); second radiomedial vein (r-m) of fore wing present; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein); vein 2-1A of fore wing more or less shortened ... Mononeuron Fischer, 1981
  • Propodeal carinated areas completely absent; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; vein 2-1A of fore wing not shortened... Micrommatus Marsh, 1993

167(165).
  • Occipital carina meeting hypostomal carina; metapleuron glabrous or with a very sparse/rare discret pubescence; apical maxillary palpomere modified and shorter than the previous; scutellum with (fine) lateral carinae ... Preembobracon Quicke & Butcher, 2015
  • Occipital carina not meeting hypostomal carina; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); apical maxillary palpomere not modified and as long as or longer than the previous; scutellum without lateral carinae... Labania Hedqvist, 1963

168(158).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 169
  • Propodeal carinated areas completely absent... 180

169(168).
  • Occipital carina meeting hypostomal carina ... 170
  • Occipital carina not meeting hypostomal carina... 177

170(169).
  • Propodeum completely or partly (micro)sculptured (e.g. Fig. 2E) ... 171
  • Propodeum almost entirely smooth (visible in the carinated areola)... 176

171(170).
  • Third labial palp segment as long as the second ... 172
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... Araucania Marsh, 1993

172(171).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... 173
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... 175

173(172).
  • Scape less than two times longer than maximally wide; scape shorter than first flagellomere ... 174
  • Scape two or more times longer than maximally wide; scape (sub)equal to first flagellomere... Whartonius Marsh, 1993

174(173).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein); frons not excavated (flat or convex); shape of second metasomal suture distinctly curved laterally; bar posterior to valvillus or valvillar zone present ... Donquickeia Marsh, 1997
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein); frons excavated (concave, sometimes weakly); shape of second metasomal suture straight or evenly curved (e.g. Fig. 7F); bar posterior to valvillus or valvillar zone absent... Allorhogas Gahan, 1912

175(172).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width ... Spathiospilus Marsh, 1999
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width... Spathius Nees, 1918

176(170).
  • Recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C) ... Waitaca Marsh, 1993
  • Recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D)... Lissodoryctes Marsh, 2002

177(169).
  • Third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G) ... 178
  • Third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... Tripteroides Marsh, 2002
  • Third tergite with a posterior smooth area bordered by a sinuous carina with small setae (also on T4-5)... Parsteres Gadelha, Nunes & De Oliveira, 2016

178(177).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width ... Guaygata Marsh, 1993
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width... 179

179(178).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)) ... Spathius Nees, 1918
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron glabrous or with a very sparse/rare discret pubescence... Glaucia Braga, Barbalho & Penteado-Dias, 2002

180(168).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); head with long pilosity somewhere (face, vertex, fully,...) ... Cyphodoryctes Marsh, 1997
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); head with sparse pilosity (short or long, sometimes in punctuations)... 181

181(180).
  • Scape less than two times longer than maximally wide; hind wing without the recurrent vein (m-cu); hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D); dorsope of first metasomal tergite absent or indistinct ... Subcurtisella Roman, 1924
  • Scape two or more times longer than maximally wide; hind wing with the recurrent vein (m-cu); hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C); dorsope of first metasomal tergite present and more or less distinct... Whartonius Marsh, 1993

182(157).
  • Ovipositor nodus without visible node (but lower valves serrate) ... 183
  • Ovipositor nodus single (e.g. Quicke et al., 1992a: figs 9-11)... Tarasco Marsh, 1993
  • Ovipositor nodus double, with a second node weakly developed (e.g. Quicke et al., 1992a: fig. 12)... 184
  • Ovipositor nodus double (or more as Cryptodoryctes), with both nodus well developed (e.g. Quicke et al., 1992a: figs 1-8)... 185

183(182).
  • Scape less than two times longer than maximally wide; mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); propodeum without lateral tubercles ... Heterospathius Barbalho & Penteado-Dias, 1999
  • Scape two or more times longer than maximally wide; mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); propodeum with lateral tubercles (as high as large at base)... Caingangia Marsh, 1993

184(182).
  • Ocelar triangle equilateral; mesoscutum surface of median and lateral lobes smooth and shining; metapleuron glabrous or with a very sparse/rare discret pubescence; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated ... Fritziella Marsh, 1993
  • Ocelar triangle with base larger than its sides; mesoscutum surface of median and lateral lobes more or less sculptured; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); second radiomedial vein (r-m) of fore wing present... Amazondoryctes Barbalho & Penteado-Dias, 1999

185(182).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present) ... 186
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent)... 189

186(185).
  • Labial palpi four-segmented; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); propodeal bridge between abdominal and coxal foramina absent; second radiomedial vein (r-m) of fore wing present ... 187
  • Labial palpi three-segmented; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); propodeal bridge between abdominal and coxal foramina present; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated... 188

187(186).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; ocelar triangle equilateral; mesoscutum surface of median and lateral lobes more or less sculptured; metapleuron glabrous or with a very sparse/rare discret pubescence ... Acanthorhogas Szepligeti, 1906
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified); ocelar triangle with base larger than its sides; mesoscutum surface of median and lateral lobes smooth and shining; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body))... Tripteria Enderlein, 1912

188(186).
  • Apical lobe of scape absent; notauli not meeting each other; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); maxillary palpi five-segmented ... Aivalykus Nixon, 1938
  • Apical lobe of scape present (apically or sub-apically); notauli meeting a sculptured area before scutelar sulcus; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); maxillary palpi six-segmented... Achterbergia Marsh, 1993

189(185).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E); first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused ... 190
  • Propodeal carinated areas completely absent; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated... 191

190(189).
  • Notauli meeting before the scutelar sulcus; scape (sub)equal to first flagellomere; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Leptodoryctes Barbalho & Penteado-Dias, 1999
  • Notauli meeting at the scutelar sulcus; scape shorter than first flagellomere; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Dapsilitas Braet, Barbalhoa & van Achterberg, 2003

191(189).
  • Occipital carina meeting hypostomal carina; metanotum with a short and acute erected spine medially; fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G); petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width ... Notiospathius Matthews & Marsh, 1973
  • Occipital carina not meeting hypostomal carina; metanotum without any protuberances or spines; fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67); petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width... 192

192(191).
  • Vertex not striate; pronotum dorsally without modifications; propodeum with lateral tubercles (as high as large at base); second radiomedial vein (r-m) of fore wing present ... Hemidoryctes Belokobylskij, 1993
  • Vertex striate; pronotum with convex lobe; propodeum without lateral tubercles; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated... Monolexis Foerster, 1862

193(156).
  • Hind wing with the nervellus (cu-a) vein inclivous (e.g. Figs 3E, 4B)(inclinated to base of wing) ... 194
  • Hind wing with the nervellus (cu-a) vein reclivous (e.g. Figs 3E, 4B)(inclinated to apex of wing)... 199
  • Hind wing without the nervellus (cu-a) vein (e.g. Fig. 4A, E)... Psenobolus Reinhard, 1885 (female and winged form)

194(193).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; sternaulus (precoxal sulcus) present; head with sparse pilosity (short or long, sometimes in punctuations) ... 195
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified); sternaulus (precoxal sulcus) absent; head with long pilosity somewhere (face, vertex, fully,...)... Jataiella Barbalho & Penteado-Dias, 1999

195(194).
  • Second radiomedial vein (r-m) of fore wing present; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; medial (basal) cell of hind wing closed (e.g. Fig. 3E) ... 196
  • Second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; medial (basal) cell of hind wing widely open antero-distally (vein SC+R absent)(e.g. Fig. 4G)... Canchim Barbalho & Penteado-Dias, 1999

196(195).
  • Propodeum without lateral tubercles; scape as long as ventrally than dorsally ... 197
  • Propodeum with lateral tubercles (as high as large at base); scape longer dorsally than ventrally... 198

197(196).
  • Scape less than two times longer than maximally wide; mesoscutum surface of median and lateral lobes smooth and shining; scape shorter than first flagellomere; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Evaniodes Szépligeti, 1901
  • Scape two or more times longer than maximally wide; mesoscutum surface of median and lateral lobes more or less sculptured; scape (sub)equal to first flagellomere; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Heredius Marsh, 2002

198(196).
  • Third labial palp segment as long as the second; frons without lateral carinae or grooves; metanotum with a short and acute erected spine medially; propodeal carinated areas completely absent ... Notiospathius Matthews & Marsh, 1973
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113); frons with some lateral carinae or grooves along the eyes; metanotum without any protuberances or spines; propodeal carinated areas present at least basally (e.g. Fig. 2E)... Monitoriella Hedqvist, 1963

199(193).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; occipital carina meeting hypostomal carina; pronotum short (nearly invisible in dorsal view); scape less than two times longer than maximally wide ... Heerz Marsh, 1993
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified); occipital carina not meeting hypostomal carina; pronotum normal to long (visible in dorsal view); scape two or more times longer than maximally wide... Lissopius Marsh, 2002

200(155).
  • Scape (sub)equal to first flagellomere ... 201
  • Scape shorter than first flagellomere... 203

201(200).
  • Pronotum short (nearly invisible in dorsal view); propodeum with lateral tubercles (as high as large at base); vein r of fore wing arising distinctly before middle of pterostigma; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated ... 202
  • Pronotum normal to long (visible in dorsal view); propodeum without lateral tubercles; vein r of fore wing arising submedially from pterostigma; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused... Neoheterospilus (Harpoheterospilus) Belokobylskij, 2006

202(201).
  • Scape less than two times longer than maximally wide; notauli not meeting each other; second radiomedial vein (r-m) of fore wing present; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein) ... Cryptontsira Belokobylskij, 2008
  • Scape two or more times longer than maximally wide; notauli meeting a sculptured area before scutelar sulcus; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Hecabolus Curtis, 1837

203(200).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 204
  • Propodeal carinated areas completely absent... Masonius Marsh, 1993

204(203).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64); raised ridge between antennae present; mid tibia with spines on antero dorsal side ... Bolivar Zaldívar-Riverón & Rodríguez-Jiménez, 2013
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63); raised ridge between antennae absent; mid tibia without spines on antero dorsal side... Spathius Nees, 1918

205(154).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base ... 206
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified)... Whitfieldiellus Marsh, 1997
  • Recurrent vein (m-cu) of hind wing strongly curved to apex of wing (fully sclerified usually)... 214

206(205).
  • Brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present) ... 207
  • Brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent)... 210

207(206).
  • Nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F) ... 208
  • Nervellus (cu-a) of fore wing interstitial... Liobracon Szépligeti, 1901
  • Nervellus (cu-a) of fore wing antefurcal (e.g. Fig. 4E)... Hybodoryctes Szépligeti, 1906

208(207).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... Trigonophasmus Enderlein, 1912
  • Fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B)... 209
  • Fourth-sixth metasomal tergites of males strongly sclerotised, distinctly convex, entirely sculptured, sixth tergite enlarged, covered following tergites... Pedinotus Szépligeti, 1902

209(208).
  • Mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); metapleuron glabrous or with a very sparse/rare discret pubescence; propodeal carinated areas completely absent ... Gymnobracon Szépligeti, 1902
  • Mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); propodeal carinated areas present at least basally (e.g. Fig. 2E)... Osmophila Szépligeti, 1902

210(206).
  • Petiole median carinae very short (less than or equal to the third of petiole length) ... Vanderentiellus Marsh, 2002
  • Petiole median carinae reaching, or nearly, the half length of the petiole... 211
  • Petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... 212

211(210).
  • Maxillary palpi six-segmented; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); hind wing with the nervellus (cu-a) vein inclivous (e.g. Figs 3E, 4B)(inclinated to base of wing) ... Sericobracon Shaw & Edgerly, 1985
  • Maxillary palpi five-segmented; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); hind wing with the nervellus (cu-a) vein reclivous (e.g. Figs 3E, 4B)(inclinated to apex of wing)... Heerz Marsh, 1993

212(210).
  • Vein r of fore wing arising distinctly before middle of pterostigma; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; first flagellar segment longer than second (e.g. Fig. 1H, I); hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K) ... 213
  • Vein r of fore wing arising submedially from pterostigma; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; first flagellar segment (sub)equal to second (e.g. Fig. 1L); hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C)... Pioscelus Muesebeck & Walkley, 1951

213(212).
  • Scape less than two times longer than maximally wide; scutellum flat; metanotum without any protuberances or spines; metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)) ... Glyptocolastes Ashmead, 1900
  • Scape two or more times longer than maximally wide; scutellum convex (e.g. swollen, conical, sharply raised); metanotum with a short (obtuse) median tubercle; metapleuron glabrous or with a very sparse/rare discret pubescence... Hecabolus Curtis, 1837

214(205).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E); scape shorter than first flagellomere; hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D); mid tibia with spines on antero dorsal side ... Odontobracon Cameron, 1887
  • Propodeal carinated areas completely absent; scape (sub)equal to first flagellomere; hind coxa dorsally without teeth (e.g. Fig. 5A, B, E); mid tibia without spines on antero dorsal side... Holcobracon Cameron, 1905

215(138).
  • Occipital carina present dorsally and laterally ... 216
  • Occipital carina present laterally and interrupted dorsally... Embobracon van Achterberg, 1995
  • Occipital carina absent... 219

216(215).
  • Scutellum flat; hind wing with the nervellus (cu-a) vein inclivous (e.g. Figs 3E, 4B)(inclinated to base of wing) ... 217
  • Scutellum convex (e.g. swollen, conical, sharply raised); hind wing without the nervellus (cu-a) vein (e.g. Fig. 4A, E)... Percnobracon Kieffer & Jörgensen, 1910

217(216).
  • Mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa); propodeum with lateral tubercles (as high as large at base); third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area); first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused ... Tuberatra Gadelha, Nunes & De Oliveira, 2016
  • Mesoscutum surface of median and lateral lobes more or less sculptured; propodeum without lateral tubercles; third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G); first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated... 218

218(217).
  • Malar suture absent or indistinct; vertex not striate; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)) ... Stenocorse Marsh, 1968
  • Malar suture present; vertex striate; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron glabrous or with a very sparse/rare discret pubescence... Percnobraconoides Marsh, 1989

219(215).
  • Scape (sub)equal to first flagellomere ... Termitobracon Brues, 1923 (male)
  • Scape shorter than first flagellomere... Ypsistocerus Cushman, 1923
  • Scape longer than first flagellomere... Termitobracon Brues, 1923 (female)

220(137).
  • Occipital carina present dorsally and laterally; propodeum completely or partly (micro)sculptured (e.g. Fig. 2E) ... 221
  • Occipital carina absent; propodeum almost entirely smooth (visible in the carinated areola)... 245

221(220).
  • First and second tergites not fused (e.g. Fig. 5G-J) ... 222
  • First and second tergites fused (e.g. Belokobylskij, 2000: figs 10, 11, 23, 24; Barbalho & Penteado-Dias, 2002: fig. 4)... Iare Barbalho & Penteado-Dias, 2002

222(221).
  • Dorsope of first metasomal tergite present and more or less distinct ... 223
  • Dorsope of first metasomal tergite absent or indistinct... 241

223(222).
  • Epicnemial carina (prepectal carina) present laterally and ventrally ... 224
  • Epicnemial carina (prepectal carina) present only ventrally... Callihormius Ashmead, 1900b
  • Epicnemial carina (prepectal carina) absent... Ptesimogastroides Braet & van Achterberg, 2001

224(223).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... 225
  • Acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E)... 233
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... 239

225(224).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... 226
  • Fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B)... 231

226(225).
  • Petiole without median carinae ... Curtisella Spinola, 1853
  • Petiole with median carinae... 227

227(226).
  • Petiole median carinae reaching, or nearly, the half length of the petiole ... 228
  • Petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... 229

228(227).
  • Pronotum with convex lobe; pronotum carina(e) or keel absent or indistinct; mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Dicarinoryctes Braet & van Achterberg, 2001
  • Pronotum dorsally without modifications; pronotum carina(e) or keel present (laterally and-or dorsally); mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Panama Marsh, 1993

229(227).
  • First flagellar segment straigth; head with sparse pilosity (short or long, sometimes in punctuations) ... 230
  • First flagellar segment more or less curved; head with long pilosity somewhere (face, vertex, fully,...)... Callihormius Ashmead, 1900b

230(229).
  • Occipital carina meeting hypostomal carina; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); fore femur with more or less distinct dorsal blister; scape longer dorsally than ventrally ... Heterospilus Halliday, 1836
  • Occipital carina not meeting hypostomal carina; propodeum clearly convex; fore femur without dorsal blister; scape as long as ventrally than dorsally... Leluthia Cameron, 1887 (female)

231(225).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron) ... Shawius Marsh, 1993
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron)... 232

232(231).
  • Occipital carina meeting hypostomal carina; propodeal carinated areas present at least basally (e.g. Fig. 2E); recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C) ... Rimacollus Marsh, 2002
  • Occipital carina not meeting hypostomal carina; propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K)... Leluthia Cameron, 1887 (male)

233(224).
  • Fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G) ... 234
  • Fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67)... 235

234(233).
  • Ocelar triangle with base larger than its sides; pronotum with convex lobe; scutellum flat; propodeal bridge between abdominal and coxal foramina absent ... Janzenia Marsh, 1993
  • Ocelar triangle absent (one or all ocelli absent); pronotum dorsally without modifications; scutellum convex (e.g. swollen, conical, sharply raised); propodeal bridge between abdominal and coxal foramina present... Oroceguera Seltmann & Sharkey, 2007

235(233).
  • Pronotum with convex lobe; scape longer dorsally than ventrally ... 236
  • Pronotum dorsally without modifications; scape as long as ventrally than dorsally... 237

236(235).
  • Pronotum normal to long (visible in dorsal view); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F); vein 2-1A of fore wing not shortened ... Callihormius Ashmead, 1900b
  • Pronotum very long (in dorsal view); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); nervellus (cu-a) of fore wing interstitial; vein 2-1A of fore wing more or less shortened... Aphelopsia Marsh, 1993

237(235).
  • Maxillary palpi six-segmented; labial palpi four-segmented; scutellum flat; wings present ... 238
  • Maxillary palpi four-segmented; labial palpi three-segmented; scutellum convex (e.g. swollen, conical, sharply raised); wings absent... Ecphylopsis Ashmead, 1900

238(237).
  • Vertex not striate; notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS) ... Platydoryctes Barbalhoa & Penteado-Dias, 2000
  • Vertex striate; notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent)... Ptesimogaster Marsh, 1965

239(224).
  • Basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F); petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63); hind tibia without (comb or area) of dense setae at inner apex ... 240
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J); petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64); hind tibia with (comb or area) of dense setae at inner apex... Sharkeyella Marsh, 1993

240(239).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); propodeal carinated areas present at least basally (e.g. Fig. 2E) ... Spathius Nees, 1918
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); propodeal carinated areas completely absent... Platydoryctes Barbalhoa & Penteado-Dias, 2000

241(222).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 242
  • Propodeal carinated areas completely absent... 244

242(241).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 243
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Angelica Marsh, 2002

243(242).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body)); petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; petiole with median carinae ... Spathius Nees, 1918
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron glabrous or with a very sparse/rare discret pubescence; petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; petiole without median carinae... Curtiselloides Marsh, 2002

244(241).
  • Occipital carina meeting hypostomal carina; vertex striate; nervellus (cu-a) of fore wing interstitial; basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F) ... Lianus Gomes & Penteado-Dias, 2006
  • Occipital carina not meeting hypostomal carina; vertex not striate; nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F); basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J)... Trigonophasmus Enderlein, 1912

245(220).
  • Apical margin of scape and pedicel without dense cluster of setae; scape shorter than first flagellomere; first flagellar segment (sub)equal to second (e.g. Fig. 1L); third labial palp segment as long as the second ... Psenobolus Reinhard, 1885 (male and brachypterous form)
  • Apical margin of scape and pedicel with dense cluster of setae; scape longer than first flagellomere; first flagellar segment longer than second (e.g. Fig. 1H, I); third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... Binarea Brullé, 1846

246(135).
  • Wings present; hind femora not swollen (at least 3x longer than wide) ... Ecphylus Foerster, 1862 (New World taxa)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

247(1).
  • Epicnemial carina (prepectal carina) present laterally and ventrally ... 248
  • Epicnemial carina (prepectal carina) present only ventrally... 295
  • Epicnemial carina (prepectal carina) absent... 296

248(247).
  • Apical lobe of scape absent ... 249
  • Apical lobe of scape present (apically or sub-apically)... 294

249(248).
  • Propodeal bridge between abdominal and coxal foramina absent ... 250
  • Propodeal bridge between abdominal and coxal foramina present... 293

250(249).
  • Second and third tergites not enlarged, nor covering following tergites ... 251
  • Second and third tergites enlarged, covering the following tegites... Chelonodoryctes Belokobylskij & Quicke, 2000

251(250).
  • First and second tergites not fused (e.g. Fig. 5G-J) ... 252
  • First and second tergites fused (e.g. Belokobylskij, 2000: figs 10, 11, 23, 24; Barbalho & Penteado-Dias, 2002: fig. 4)... 292

252(251).
  • Scape less than two times longer than maximally wide ... 253
  • Scape two or more times longer than maximally wide... Spathius (Antespathius) Belokobylskij, 1995

253(252).
  • Second abscissa of costal vein in hind wing absent; metasoma of male not depressed, rather long, not specially curved ... 254
  • Second abscissa of costal vein in hind wing usually present (sclerotized); metasoma of male not depressed dorsoventrally, relatively short, curved down apically... 291

254(253).
  • Hind femur without teeth on ventral side ... 255
  • Hind femur with more than one tooth (often one or two and several smaller) on ventral side... Euscelinus Westwood, 1882

255(254).
  • Sixth tergite visible ... 256
  • Sixth tergite hidden (by the fifth tergite)... Rhaconotus Ruthe, 1854

256(255).
  • Dorsal tentorial pits near the antennal sockets absent ... 257
  • Dorsal tentorial pits near the antennal sockets present between antenal sockets and eyes... Neurocrassus Snoflak, 1945
  • Dorsal tentorial pits near the antennal sockets present between antenal sockets only... Ontsira Cameron, 1900

257(256).
  • First flagellar segment surface dorso-ventrally not compressed ... 258
  • First flagellar segment surface dorso-ventrally compressed (sometimes strongly)... 290

258(257).
  • Dorsope of first metasomal tergite present and more or less distinct ... 259
  • Dorsope of first metasomal tergite absent or indistinct... 288

259(258).
  • Third labial palp segment as long as the second ... 260
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... 287

260(259).
  • Petiole without median carinae ... Halycaea Cameron, 1903
  • Petiole with median carinae... 261

261(260).
  • Scape (sub)equal to first flagellomere ... 262
  • Scape shorter than first flagellomere... 265
  • Scape longer than first flagellomere... Ipodoryctes Granger, 1949

262(261).
  • Propodeum without lateral tubercles; petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B); scape as long as ventrally than dorsally ... 263
  • Propodeum with lateral tubercles (as high as large at base); petiole median carinae reaching, or nearly, the half length of the petiole; fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly; scape longer dorsally than ventrally... 264

263(262).
  • Pronotum with convex lobe; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; median lobe of mesoscutum with lateral corners anteriorly angled, toothed or (very) acute; mesoscutum surface of median and lateral lobes smooth and shining ... Neoheterospilus (Neoheterospilus) Belokobylskij, 2006
  • Pronotum dorsally without modifications; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; median lobe of mesoscutum rounded antero-laterally; mesoscutum surface of median and lateral lobes more or less sculptured... Ipodoryctes Granger, 1949

264(262).
  • Eyes not emarginated; vein r of fore wing arising distinctly after(behind) middle of pterostigma; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Spathiomorpha Tobias, 1976
  • Eyes more or less emarginated on their inner sides; vein r of fore wing arising distinctly before middle of pterostigma; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Cryptontsira Belokobylskij, 2008

265(261).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence ... 266
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body))... 284

266(265).
  • Mesoscutum surface of median and lateral lobes smooth and shining ... 267
  • Mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa)... Heterospilus Halliday, 1836
  • Mesoscutum surface of median and lateral lobes more or less sculptured... 271

267(266).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F); second metasomal suture present (e.g. Fig. 7C, D, F) ... 268
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); second metasomal suture indistinct, to largely or entirely absent (e.g. Figs 6B, E, 7E)... 270

268(267).
  • Recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); nervellus (cu-a) of fore wing interstitial; scape as long as ventrally than dorsally; mid femora with more or less distinct dorsal blister ... Hypodoryctes Kokujev, 1900
  • Recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F); scape longer dorsally than ventrally; mid femora without dorsal blister... 269

269(268).
  • Pronotum short (nearly invisible in dorsal view); pronotum carina(e) or keel absent or indistinct; brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); vein 2-1A of fore wing more or less shortened ... Parallorhogas Marsh, 1993
  • Pronotum normal to long (visible in dorsal view); pronotum carina(e) or keel present (laterally and-or dorsally); brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); vein 2-1A of fore wing not shortened... Doryctes (Doryctes + Neodoryctes) Haliday, 1836

270(267).
  • Ocelar triangle equilateral; median lobe of mesoscutum with lateral corners anteriorly angled, toothed or (very) acute; scutellum convex (e.g. swollen, conical, sharply raised); propodeum with lateral long spines (higher than its base) ... Ryukyuspathius Belokobylskij, 2008
  • Ocelar triangle with base larger than its sides; median lobe of mesoscutum rounded antero-laterally; scutellum flat; propodeum with lateral tubercles (as high as large at base)... Spathius (Ambispathius) Belokobylskij, 1995

271(266).
  • Basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F) ... 272
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J)... 282

272(271).
  • Fore femur with more or less distinct dorsal blister; mid femora with more or less distinct dorsal blister ... 273
  • Fore femur without dorsal blister; mid femora without dorsal blister... 276

273(272).
  • Frons without lateral carinae or grooves; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); propodeum completely or partly (micro)sculptured (e.g. Fig. 2E); hind tibia without (comb or area) of dense setae at inner apex ... 274
  • Frons with some lateral carinae or grooves along the eyes; propodeum clearly convex; propodeum almost entirely smooth (visible in the carinated areola); hind tibia with (comb or area) of dense setae at inner apex... Asiaontsira Belokobylskij, Pu and Chen, 2013

274(273).
  • Mid tibia without spines on antero dorsal side; male genitalia with basal lobe of basal ring (e.g. Fig. 7B, G) ... 275
  • Mid tibia with spines on antero dorsal side; male genitalia without basal lobe of basal ring (e.g. Fig. 8A, C)... Rhacontsira Belokobylskij, 1998

275(274).
  • Hind femora with more or less distinct dorsal blister; comprising two parts; bar posterior to valvillus or valvillar zone absent; bars anterior to valvillus or valvillar zone absent ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts; bar posterior to valvillus or valvillar zone present; bars anterior to valvillus or valvillar zone present... Ontsira Cameron, 1900

276(272).
  • Pronotum with convex lobe ... 277
  • Pronotum dorsally without modifications... 278

277(276).
  • Vertex not striate; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; petiole median carinae reaching, or nearly, the half length of the petiole; scape more or less flattened laterally (somewhere on it) ... Caenophanes Foerster, 1862
  • Vertex striate; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; petiole median carinae very short (less than or equal to the third of petiole length); scape not flattened laterally... Monolexis Foerster, 1862

278(276).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 279
  • Propodeal carinated areas completely absent... 280

279(278).
  • Pronotum short (nearly invisible in dorsal view); acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F); head with long pilosity somewhere (face, vertex, fully,...); mid tibia without spines on antero dorsal side ... Dendrosotinus Telenga, 1941
  • Pronotum normal to long (visible in dorsal view); acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); head with sparse pilosity (short or long, sometimes in punctuations); mid tibia with spines on antero dorsal side... Spathius (Ambispathius) Belokobylskij, 1995

280(278).
  • Propodeum without lateral tubercles; vein 2-1A of fore wing not shortened; petiole median carinae reaching, or nearly, the half length of the petiole; hind tibia without (comb or area) of dense setae at inner apex ... 281
  • Propodeum with lateral tubercles (as high as large at base); vein 2-1A of fore wing more or less shortened; petiole median carinae very short (less than or equal to the third of petiole length); hind tibia with (comb or area) of dense setae at inner apex... Hemidoryctes Belokobylskij, 1993

281(280).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); second radiomedial vein (r-m) of fore wing present; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Dendrosoter Wesmael, 1838
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Hecabalodes Wilkinson, 1929

282(271).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Eodendrus Belokobylskij, 1998
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein)... Hecabalodes Wilkinson, 1929
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein)... 283

283(282).
  • Clypeus without lower flange; pronotum very long (in dorsal view); vein 2-1A of fore wing more or less shortened; hind wing without the recurrent vein (m-cu) ... Spathiostenus Belokobylskij, 1992
  • Clypeus with lower flange more or less developped; pronotum normal to long (visible in dorsal view); vein 2-1A of fore wing not shortened; hind wing with the recurrent vein (m-cu)... Polystenus Förster, 1862

284(265).
  • Petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width ... Pedinotus (Eopedinotus) Belokobylskij, 2004
  • Petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width... 285

285(284).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); propodeal carinated areas present at least basally (e.g. Fig. 2E); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 286
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); propodeal carinated areas completely absent; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Polystenus Förster, 1862

286(285).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F); fore femur without dorsal blister ... Spathius Nees, 1918
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F); basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J); fore femur with more or less distinct dorsal blister... Hecabolomorpha Belokobylskij & Chen, 2006

287(259).
  • Recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); second radiomedial vein (r-m) of fore wing with wide bulla (e.g. Fig. 3d); hind coxa dorsally without teeth (e.g. Fig. 5A, B, E); scape as long as ventrally than dorsally ... Holcobracon Cameron, 1905
  • Recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); second radiomedial vein (r-m) of fore wing largely tubular (e.g. Fig. 4H; Marsh, 1997: figs 16, 18, 30); hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D); scape longer dorsally than ventrally... Zombrus Marshall, 1897

288(258).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... 289
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Parana Nixon, 1943

289(288).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... Halycaea Cameron, 1903
  • Acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E)... Platyspathius Viereck, 1911
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Spathius Nees, 1918

290(257).
  • Pronotum short (nearly invisible in dorsal view); pronotum dorsally without modifications; pronotum carina(e) or keel present (laterally and-or dorsally); propodeal carinated areas present at least basally (e.g. Fig. 2E) ... Dendrosotinus Telenga, 1941
  • Pronotum very long (in dorsal view); pronotum with convex lobe; pronotum carina(e) or keel absent or indistinct; propodeal carinated areas completely absent... Doryctophasmus Enderlein, 1912

291(253).
  • Wings present; hind femora not swollen (at least 3x longer than wide); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... Ecphylus Foerster, 1862 (winged forms)
  • Wings micropterous to scalelike pads; hind femora swollen, at most 2.6x longer than wide; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Ecphylus Foerster, 1862 (apterous or brachypterous forms)

292(251).
  • First flagellar segment longer than second (e.g. Fig. 1H, I); sixth tergite hidden (by the fifth tergite); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein); fore tibial spines more or less numerous and dispersed (e.g. Fig. 5G) ... Mimipodoryctes Belokobylskij, 2000
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); sixth tergite visible; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines usually few and forming a single row (e.g. Marsh, 1997: figs 65-67)... Arhaconotus Belokobylskij, 2000

293(249).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... Aivalykus Nixon, 1938
  • Acrosternite of first metasomal tergite long, 0.3-0.5 as long as tergite and fused with its ventral margin anteriorly, petiole present (e.g. Fig. 6D, E)... Leptospathius Szépligeti, 1902
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Australospathius Belokobylskij, Iqbal & Austin, 2004

294(248).
  • Metanotum without any protuberances or spines; scape longer ventrally than dorsally ... Synspilus Belokobylskij & Quicke, 2000
  • Metanotum with a short and acute erected spine medially; scape longer dorsally than ventrally... Paraspathius Nixon, 1943
  • Metanotum with a short (obtuse) median tubercle; scape as long as ventrally than dorsally... Arkoola Belokobylskij, Iqbal & Austin, 2004

295(247).
  • First flagellar segment longer than second (e.g. Fig. 1H, I); occipital carina present laterally and interrupted dorsally; wings present ... Termitospathius Belokobylskij, 2002 (female)
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); occipital carina present dorsally and laterally; wings reduced or brachypterous (wings shorter than metasoma)... Termitospathius Belokobylskij, 2002 (male)

296(247).
  • Apical lobe of scape absent; scape (sub)equal to first flagellomere; first flagellar segment shorter than second; maxillary palpi five-segmented ... Ceylonspathius Belokobylskij, 2002
  • Apical lobe of scape present (apically or sub-apically); scape shorter than first flagellomere; first flagellar segment longer than second (e.g. Fig. 1H, I); maxillary palpi six-segmented... Siragra Cameron, 1907

297(1).
  • Apical lobe of scape absent ... 298
  • Apical lobe of scape present (apically or sub-apically)... 348

298(297).
  • Epicnemial carina (prepectal carina) present laterally and ventrally ... 299
  • Epicnemial carina (prepectal carina) present only ventrally... Dooliba Belokobylskij, Iqbal & Austin, 2004
  • Epicnemial carina (prepectal carina) absent... Bracodoryctes Belokobylskij & Quicke, 2000

299(298).
  • Notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14) ... 300
  • Notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86)... 347

300(299).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence ... 301
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body))... 339

301(300).
  • Dorsope of first metasomal tergite present and more or less distinct ... 302
  • Dorsope of first metasomal tergite absent or indistinct... 336

302(301).
  • Propodeal bridge between abdominal and coxal foramina absent ... 303
  • Propodeal bridge between abdominal and coxal foramina present... Fijibracon Belokobylskij, 1995

303(302).
  • Hind femur without teeth on ventral side ... 304
  • Hind femur with more than one tooth (often one or two and several smaller) on ventral side... Euscelinus Westwood, 1882

304(303).
  • First and second tergites not fused (e.g. Fig. 5G-J) ... 305
  • First and second tergites fused (e.g. Belokobylskij, 2000: figs 10, 11, 23, 24; Barbalho & Penteado-Dias, 2002: fig. 4)... Mimipodoryctes Belokobylskij, 2000

305(304).
  • Scape less than two times longer than maximally wide ... 306
  • Scape two or more times longer than maximally wide... Kauriphanes Belokobylskij & Zaldívar, 2011

306(305).
  • First flagellar segment surface dorso-ventrally not compressed ... 307
  • First flagellar segment surface dorso-ventrally compressed (sometimes strongly)... 335

307(306).
  • Median lobe of mesoscutum rounded antero-laterally ... 308
  • Median lobe of mesoscutum with lateral corners anteriorly angled, toothed or (very) acute... Neoheterospilus (Neoheterospilus) Belokobylskij, 2006

308(307).
  • Labial palpi four-segmented ... 309
  • Labial palpi three-segmented... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Labial palpi two-segmented (e.g. van Achterberg, 1995: fig. 46)... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Labial palpi one-segmented... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Labial palpi absent (e.g. van Achterberg, 1995: fig. 56)... Nipponecphylus Belokobylskij & Konishi, 2001 (males)

309(308).
  • Third labial palp segment as long as the second ... 310
  • Third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113)... Zombrus Marshall, 1897

310(309).
  • Hind coxa dorsally without teeth (e.g. Fig. 5A, B, E) ... 311
  • Hind coxa with one to several teeth dorsally (e.g. Fig. 5C, D)... Echinodoryctes Belokobylskij, Iqbal & Austin, 2004

311(310).
  • Scape not flattened laterally ... 312
  • Scape more or less flattened laterally (somewhere on it)... 334

312(311).
  • Basal area of second tergite enclosed by carinae or grooves absent (e.g. Fig. 6B, F) ... 313
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) (e.g. Fig. 6G-J)... 330

313(312).
  • Fore femur with more or less distinct dorsal blister ... 314
  • Fore femur without dorsal blister... 319

314(313).
  • Frons without lateral carinae or grooves; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 315
  • Frons with some lateral carinae or grooves along the eyes; propodeum clearly convex... Asiaontsira Belokobylskij, Pu and Chen, 2013

315(314).
  • Fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B) ... 316
  • Fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface) (e.g. Fig. 7B); fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F)... Ipodoryctes Granger, 1949

316(315).
  • Mid tibia without spines on antero dorsal side ... 317
  • Mid tibia with spines on antero dorsal side... Rhacontsira Belokobylskij, 1998

317(316).
  • Hind femora with more or less distinct dorsal blister; comprising two parts ... Heterospilus Halliday, 1836
  • Hind femora without dorsal blister; comprising three parts... 318

318(317).
  • Pronotum carina(e) or keel absent or indistinct; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; mid femora without dorsal blister ... Parallorhogas Marsh, 1993
  • Pronotum carina(e) or keel present (laterally and-or dorsally); mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); mid femora with more or less distinct dorsal blister... Ontsira Cameron, 1900

319(313).
  • Pronotum short (nearly invisible in dorsal view) ... 320
  • Pronotum normal to long (visible in dorsal view)... 326
  • Pronotum very long (in dorsal view)... Spathiomorpha Tobias, 1976

320(319).
  • Acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F); propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); propodeum completely or partly (micro)sculptured (e.g. Fig. 2E) ... 321
  • Acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B); propodeum clearly convex; propodeum almost entirely smooth (visible in the carinated areola)... Fijispathius Belokobylskij, Iqbal & Austin, 2004

321(320).
  • Scape (sub)equal to first flagellomere ... 322
  • Scape shorter than first flagellomere... 323

322(321).
  • Eyes not emarginated; pronotum carina(e) or keel absent or indistinct; notauli meeting a sculptured area before scutelar sulcus; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron) ... Rhoptrocentrus Marshall, 1897
  • Eyes more or less emarginated on their inner sides; pronotum carina(e) or keel present (laterally and-or dorsally); notauli not meeting each other; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron)... Cryptontsira Belokobylskij, 2008

323(321).
  • Pronotum carina(e) or keel absent or indistinct; first flagellar segment straigth; hind tibia with (comb or area) of dense setae at inner apex ... 324
  • Pronotum carina(e) or keel present (laterally and-or dorsally); first flagellar segment more or less curved; hind tibia without (comb or area) of dense setae at inner apex... 325

324(323).
  • Vertex not striate; pronotum dorsally without modifications; propodeum with lateral tubercles (as high as large at base); second radiomedial vein (r-m) of fore wing present ... Hemidoryctes Belokobylskij, 1993
  • Vertex striate; pronotum with convex lobe; propodeum without lateral tubercles; second radiomedial vein (r-m) of fore wing absent with veins 2M and 3RSa,b widely separated... Monolexis Foerster, 1862

325(323).
  • Scutellum without lateral carinae; propodeal carinated areas completely absent; fourth-sixth metasomal tergites of males submarginal lateral carinae present (e.g. Fig. 7E); ovipositor nodus double (or more as Cryptodoryctes), with both nodus well developed (e.g. Quicke et al., 1992a: figs 1-8) ... Dendrosoter Wesmael, 1838
  • Scutellum with (fine) lateral carinae; propodeal carinated areas present at least basally (e.g. Fig. 2E); fourth-sixth metasomal tergites of males simple, submarginal lateral carinae absent; ovipositor nodus double, with a second node weakly developed (e.g. Quicke et al., 1992a: fig. 12)... Dendrosotinus Telenga, 1941

326(319).
  • Frons without lateral carinae or grooves; frons not excavated (flat or convex); hind femora without dorsal blister; second tergite without apical lenticulate area (e.g. Fig. 6B, E) ... 327
  • Frons with some lateral carinae or grooves along the eyes; frons excavated (concave, sometimes weakly); hind femora with more or less distinct dorsal blister; second tergite with apical lenticulate area (e.g. Fig. 6F, I) (often formed by the meeting of second suture and groove on third tergite when present)... Neorhaconotus Belokobylskij, Iqbal & Austin, 2004

327(326).
  • Scutellum without lateral carinae; spiral stucture of venom reservoir normal; bar posterior to valvillus or valvillar zone absent ... 328
  • Scutellum with (fine) lateral carinae; spiral stucture of venom reservoir posteriorly much courser than anteriorly; bar posterior to valvillus or valvillar zone present... 329

328(327).
  • Mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); brachial (1st subdiscal) cell of fore wing distally closed (brachial vein 2cu-a present); hind wing male without a stigma-like enlargement (e.g. Fig. 3A, C) ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); brachial (1st subdiscal) cell of fore wing open (brachial vein 2cu-a absent); hind wing male with a stigma-like enlargement in distal part of costal (1-SC + R) vein, without incurved marginal parts (e.g. Fig. 2H-K)... Leluthia Cameron, 1887 (female)

329(327).
  • Scutellum flat; petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole); scape shorter than first flagellomere; basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64) ... Spathius (Ambispathius) Belokobylskij, 1995
  • Scutellum convex (e.g. swollen, conical, sharply raised); petiole median carinae reaching, or nearly, the half length of the petiole; scape (sub)equal to first flagellomere; basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63)... Spathiomorpha Tobias, 1976

330(312).
  • Ovipositor nodus single (e.g. Quicke et al., 1992a: figs 9-11) ... Ontsirospathius Belokobylskij, Iqbal & Austin, 2004
  • Ovipositor nodus double, with a second node weakly developed (e.g. Quicke et al., 1992a: fig. 12)... 331
  • Ovipositor nodus double (or more as Cryptodoryctes), with both nodus well developed (e.g. Quicke et al., 1992a: figs 1-8)... 333

331(330).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; mesoscutum surface of median and lateral lobes smooth and shining; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M + CU more than 1times, often 1.5-2.0 times, as long as second abscissa (1-M)(e.g. Fig. 3A, C); petiole median carinae reaching, or nearly, the half length of the petiole ... Doryctes (Plyctes) Fischer, 1981
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; mesoscutum surface of median and lateral lobes more or less sculptured; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M + CU 0.2-1.0 times as long as second abscissa (1-M) (e.g. Fig. 3B, D); petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... 332

332(331).
  • Eyes not emarginated; ocelar triangle equilateral; pronotum with convex lobe; propodeal carinated areas completely absent ... Spathiostenus Belokobylskij, 1992
  • Eyes more or less emarginated on their inner sides; ocelar triangle with base larger than its sides; pronotum dorsally without modifications; propodeal carinated areas present at least basally (e.g. Fig. 2E)... Ipodoryctes Granger, 1949

333(330).
  • Recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); petiole median carinae very short (less than or equal to the third of petiole length) ... Eodendrus Belokobylskij, 1998
  • Recurrent vein (m-cu) of fore wing interstitial (arising directly in line with 2RS); petiole median carinae more than the half length of petiole (sometimes reaching the apex of petiole)... Leluthia Cameron, 1887 (male)
  • Recurrent vein (m-cu) of fore wing antefurcal (arising basad with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); petiole median carinae reaching, or nearly, the half length of the petiole... Doryctes (Doryctes + Neodoryctes) Haliday, 1836

334(311).
  • Pronotum with convex lobe; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; hind femora not swollen (at least 3x longer than wide); petiole median carinae reaching, or nearly, the half length of the petiole ... Caenophanes Foerster, 1862
  • Pronotum dorsally without modifications; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; hind femora swollen, at most 2.6x longer than wide; petiole median carinae very short (less than or equal to the third of petiole length)... Hemidoryctes Belokobylskij, 1993

335(306).
  • Pronotum short (nearly invisible in dorsal view) ... Dendrosotinus Telenga, 1941
  • Pronotum normal to long (visible in dorsal view)... Sonanus Belokobylskij & Konishi, 2001
  • Pronotum very long (in dorsal view)... Doryctophasmus Enderlein, 1912

336(301).
  • Recurrent vein (m-cu) of hind wing perpendicular or curved towards wing base; propodeal bridge between abdominal and coxal foramina absent; pronotum normal to long (visible in dorsal view); hind wing without transverse vein (r) of radial cell ... 337
  • Recurrent vein (m-cu) of hind wing more or less curved apically to apex of wing (often vein not sclerified); propodeal bridge between abdominal and coxal foramina present; pronotum very long (in dorsal view); hind wing with a transverse vein (r) of radial cell(e.g. Marsh, 1997: fig. 12)... Leptospathius Szépligeti, 1902

337(336).
  • Mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of 2-1A vein); scape (sub)equal to first flagellomere ... Pseudospathius Szepligeti, 1902
  • Mesoscutum surface of median and lateral lobes more or less sculptured; scutellum convex (e.g. swollen, conical, sharply raised); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scape shorter than first flagellomere... 338

338(337).
  • Vertex not striate; pronotum with convex lobe; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Platyspathius Viereck, 1911
  • Vertex striate; pronotum dorsally without modifications; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Spathius (Stenophasmus) Smith, 1858

339(300).
  • Propodeal carinated areas present at least basally (e.g. Fig. 2E) ... 340
  • Propodeal carinated areas completely absent... 345

340(339).
  • Third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G) ... 341
  • Third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... 344

341(340).
  • First and second tergites not fused (e.g. Fig. 5G-J); sixth tergite visible; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B); second tergite without apical lenticulate area (e.g. Fig. 6B, E) ... 342
  • First and second tergites fused (e.g. Belokobylskij, 2000: figs 10, 11, 23, 24; Barbalho & Penteado-Dias, 2002: fig. 4); sixth tergite hidden (by the fifth tergite); fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F); second tergite with apical lenticulate area (e.g. Fig. 6F, I) (often formed by the meeting of second suture and groove on third tergite when present)... Mimipodoryctes Belokobylskij, 2000

342(341).
  • Pronotum short (nearly invisible in dorsal view); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); petiole without median carinae; acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... 343
  • Pronotum normal to long (visible in dorsal view); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); petiole with median carinae; acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Spathius Nees, 1918

343(342).
  • Fore femur with more or less distinct dorsal blister; mid femora with more or less distinct dorsal blister ... genD female
  • Fore femur without dorsal blister; mid femora without dorsal blister... genD male

344(340).
  • First flagellar segment longer than second (e.g. Fig. 1H, I); malar suture present; pronotum short (nearly invisible in dorsal view); metanotum with a short (obtuse) median tubercle ... Pirramurra Belokobylskij, Iqbal & Austin, 2004
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); malar suture absent or indistinct; pronotum normal to long (visible in dorsal view); metanotum without any protuberances or spines... Arhaconotus Belokobylskij, 2000

345(339).
  • Scape (sub)equal to first flagellomere; third labial palp segment distinctly shorter than second (e.g. Belokobylskij & Quicke, 2000: fig. 113); pronotum normal to long (visible in dorsal view); nervellus (cu-a) of fore wing interstitial ... Acanthodoryctes Turner, 1918
  • Scape shorter than first flagellomere; third labial palp segment as long as the second; pronotum short (nearly invisible in dorsal view); nervellus (cu-a) of fore wing postfurcal (e.g. Fig. 3A-F)... 346

346(345).
  • Eyes not emarginated; vertex not striate; pronotum with convex lobe; petiole without median carinae ... Halycaea Cameron, 1903
  • Eyes more or less emarginated on their inner sides; vertex striate; pronotum with pointed spines or tubercles; petiole with median carinae... Ceratodoryctes Belokobylskij, Iqbal & Austin, 2004

347(299).
  • Malar suture absent or indistinct; occipital carina present laterally and interrupted dorsally; pronotum dorsally without modifications; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum ... Doryctopsis Belokobylskij, Iqbal & Austin, 2004
  • Malar suture present; occipital carina absent; pronotum with convex lobe; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum... Liodoryctes Szépligeti, 1906

348(297).
  • Pronotum with convex lobe ... 349
  • Pronotum dorsally without modifications... 350
  • Pronotum with pointed spines or tubercles... Antidoryctes Belokobylskij & Quicke, 2000

349(348).
  • Scape less than two times longer than maximally wide; scape shorter than first flagellomere; labial palpi four-segmented; malar suture absent or indistinct ... Siragra Cameron, 1907
  • Scape two or more times longer than maximally wide; scape longer than first flagellomere; labial palpi three-segmented; malar suture present... Papuaontsira Belokobylskij, 2016

350(348).
  • Fore wing hyaline ... Jarra Marsh & Austin, 1994
  • Fore wing banded... Gurburra Belokobylskij, Iqbal & Austin, 2004
  • Fore wing more or less infuscate (sometimes with pale spots inside)... 351

351(350).
  • Metanotum without any protuberances or spines; frons with some lateral carinae or grooves along the eyes; scape more or less flattened laterally (somewhere on it); scape with basal constriction ... Pseudosyngaster Belokobylskij, Iqbal & Austin, 2004 (female)
  • Metanotum with a short and acute erected spine medially; frons without lateral carinae or grooves; scape not flattened laterally; scape without basal constriction... 352

352(351).
  • Scape shorter than first flagellomere; malar suture present; occipital carina not meeting hypostomal carina; vertex striate ... Paraspathius Nixon, 1943
  • Scape longer than first flagellomere; malar suture absent or indistinct; occipital carina meeting hypostomal carina; vertex not striate... Syngaster Brullé, 1846

353(1).
  • Scape less than two times longer than maximally wide; occipital carina present dorsally and laterally ... 354
  • Scape two or more times longer than maximally wide; occipital carina absent... Sisupala Nixon, 1943

354(353).
  • Labial palpi four-segmented ... 355
  • Labial palpi three-segmented... 367

355(354).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence ... 356
  • Metapleuron with clear pubescence (setae longs, sparse or numerous, sometimes dense pubescence (in this case often present on other part of body))... 364

356(355).
  • Pronotum carina(e) or keel absent or indistinct ... 357
  • Pronotum carina(e) or keel present (laterally and-or dorsally)... 359

357(356).
  • First flagellar segment surface dorso-ventrally not compressed; scape not flattened laterally; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of 2-1A vein); hind femora not swollen (at least 3x longer than wide) ... 358
  • First flagellar segment surface dorso-ventrally compressed (sometimes strongly); scape more or less flattened laterally (somewhere on it); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; hind femora swollen, at most 2.6x longer than wide... Euscelinus Westwood, 1882

358(357).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated) ... Monolexis Foerster, 1862
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully)... Parallorhogas Marsh, 1993

359(356).
  • Petiole without median carinae; dorsope of first metasomal tergite absent or indistinct ... Spathius (Stenophasmus) Smith, 1858
  • Petiole with median carinae; dorsope of first metasomal tergite present and more or less distinct... 360

360(359).
  • Scape (sub)equal to first flagellomere; frons excavated (concave, sometimes weakly); head with long pilosity somewhere (face, vertex, fully,...) ... Cryptontsira Belokobylskij, 2008
  • Scape shorter than first flagellomere; frons not excavated (flat or convex); head with sparse pilosity (short or long, sometimes in punctuations)... 361

361(360).
  • Dorsal spines of hind tibia absent (e.g. Fig. 5H, J); propodeum completely or partly (micro)sculptured (e.g. Fig. 2E); basoventral tooth of hind coxa present (sometimes very small) (e.g. Fig. 5A; Marsh, 1997: fig. 63) ... 362
  • Dorsal spines of hind tibia present (e.g. Fig. 5I); propodeum almost entirely smooth (visible in the carinated areola); basoventral tooth of hind coxa absent (e.g. Fig. 5B-D; Marsh, 1997: fig. 64)... Fijispathius Belokobylskij, Iqbal & Austin, 2004

362(361).
  • Sixth tergite visible; hind tibia without (comb or area) of dense setae at inner apex; fifth or sixth metasomal tergites entirely smooth (e.g. Fig. 7A, B); male genitalia with basal lobe of basal ring (e.g. Fig. 7B, G) ... 363
  • Sixth tergite hidden (by the fifth tergite); hind tibia with (comb or area) of dense setae at inner apex; fifth or sixth metasomal tergites sculptured at least basally (e.g. Fig. 7C, D, F); male genitalia without basal lobe of basal ring (e.g. Fig. 8A, C)... Rhaconotus Ruthe, 1854

363(362).
  • Mesoscutum surface of median and lateral lobes smooth and shining; fore femur without dorsal blister; mid femora without dorsal blister; comprising two parts ... Doryctes (Doryctes + Neodoryctes) Haliday, 1836
  • Mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct dorsal blister; mid femora with more or less distinct dorsal blister; comprising three parts... Ontsira Cameron, 1900

364(355).
  • Scape (sub)equal to first flagellomere; notauli entirely absent (e.g. Belokobylskij & Quicke, 2000: fig. 86); propodeum clearly convex ... Stenocorse Marsh, 1968
  • Scape shorter than first flagellomere; notauli complete or at least partly present (smooth or sculptured)(e.g. Marsh, 1993: fig. 14); propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... 365

365(364).
  • Sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); third tergite without any distinct transverse narrow depression/depression (e.g. Fig. 6B, F, G); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); fore femur without dorsal blister ... 366
  • Sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); third tergite with a distinct transverse depression (furrow) (usually between its anterior third and the middle, e.g. Fig. 6H)(sometimes meeting laterally second metasomal suture and enclosing a lenticular area); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); fore femur with more or less distinct dorsal blister... Doryctinus Roman, 1910

366(365).
  • Pronotum short (nearly invisible in dorsal view); propodeal carinated areas completely absent; petiole apical width less than or equal to 1.5 times (or subequal to) minimum (often measured basally) width; acrosternite of first metasomal tergite short, nearly 0.2-0.25 as long as tergite, not fused with ventral margins of tergite, petiole absent (e.g. Fig. 6F) ... Glyptocolastes Ashmead, 1900
  • Pronotum normal to long (visible in dorsal view); propodeal carinated areas present at least basally (e.g. Fig. 2E); petiole apical width more or equal to 1.6 times (even 2 times) minimum (often measured basally) width; acrosternite of first metasomal tergite very long, 0.6-0.85 (or more) as long as tergite and entirely fused with it ventral margin, petiole present and long (Fig. 6A, B)... Spathius Nees, 1918

367(354).
  • First flagellar segment (sub)equal to second (e.g. Fig. 1L); maxillary palpi four-segmented; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; wings absent ... Ecphylopsis Ashmead, 1900
  • First flagellar segment shorter than second; maxillary palpi six-segmented; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; wings present... Fijibracon Belokobylskij, 1995
Taxonomic name: 
Cecidospathius (Doryctinae), Celereon (Doryctinae), Doryctomorpha (Doryctinae), Esterella (Doryctinae), Metaspathius (Doryctinae), Sinaodoryctes (Doryctinae), Mexiare (Doryctinae), Ecphylus (Doryctinae), Liobracon (Doryctinae), Odontobracon (Doryctinae), Pioscelus (Doryctinae), Dendrosoter (Doryctinae), Rhoptrocentrus (Doryctinae), Ptesimogaster (Doryctinae), Aivalykus (Doryctinae), Pambolidea (Doryctinae), Rhaconotus (Doryctinae), Spathius (Doryctinae), Doryctes (Doryctinae), Ontsira (Doryctinae), Allorhogas (Doryctinae), Stenocorse (Doryctinae), Parallorhogas (Doryctinae), Callihormius (Doryctinae), Heterospilus (Doryctinae), Curtisella (Doryctinae), Leluthia (Doryctinae), Hemidoryctes (Doryctinae), Acrophasmus (Doryctinae), Glyptocolastes (Doryctinae), Percnobraconoides (Doryctinae), Monolexis (Doryctinae), Aptenobracon (Doryctinae), Asiaheterospilus (Doryctinae), Nipponecphylus (Doryctinae), Cryptontsira (Doryctinae), Stenophasmus (Doryctinae), Dendrosotinus (Doryctinae), Zombrus (Doryctinae), Sonanus (Doryctinae), Eodendrus (Doryctinae), Polystenus (Doryctinae), Pareucorystes (Doryctinae), Rhacontsira (Doryctinae), Hypodoryctes (Doryctinae), Platyspathius (Doryctinae), Neurocrassus (Doryctinae), Caenophanes (Doryctinae), Spathiomorpha (Doryctinae), Euhecabolodes (Doryctinae), Hecabolomorpha (Doryctinae), Hecabolus (Doryctinae), Euscelinus (Doryctinae), Caenopachys (Doryctinae), Pseudorhaconotus (Doryctinae), Mimodoryctes (Doryctinae), Spathioplites (Doryctinae), Bulbonervus (Doryctinae), Toka (Doryctinae), Doryctophasmus (Doryctinae), Ivondrovia (Doryctinae), Leptorhaconotus (Doryctinae), Doryctoproctus (Doryctinae), Hemispathius (Doryctinae), Cryptodoryctes (Doryctinae), Gymnobracon (Doryctinae), Stephanospathius (Doryctinae), Schlettereriella (Doryctinae), Afrospathius (Doryctinae), Pararhacon (Doryctinae), Ipodoryctes (Doryctinae), Bathycentor (Doryctinae), Neoheterospilus (Doryctinae), Neoheterospilus (Doryctinae), Priosphys (Doryctinae), Odontodoryctes (Doryctinae), Paradoryctes (Doryctinae), Rinamba (Doryctinae), Pseudodoryctes (Doryctinae), Terate (Doryctinae), Grangerdoryctes (Doryctinae), Termitobracon (Doryctinae), Nervellius (Doryctinae), Psenobolus (Doryctinae), Monarea (Doryctinae), Sharkeyella (Doryctinae), Pannuceus (Doryctinae), Mononeuron (Doryctinae), Histeromeroides (Doryctinae), Megaloproctus (Doryctinae), Notiospathius (Doryctinae), Monitoriella (Doryctinae), Embobracon (Doryctinae), Holcobracon (Doryctinae), Binarea (Doryctinae), Dicarinoryctes (Doryctinae), Ptesimogastroides (Doryctinae), Rimacollus (Doryctinae), Janzenia (Doryctinae), Glaucia (Doryctinae), Amazondoryctes (Doryctinae), Heterospathius (Doryctinae), Labania (Doryctinae), Ypsistocerus (Doryctinae), Achterbergia (Doryctinae), Leptodoryctes (Doryctinae), Verae (Doryctinae), Angelica (Doryctinae), Fritziella (Doryctinae), Percnobracon (Doryctinae), Micrommatus (Doryctinae), Araucania (Doryctinae), Cyphodoryctes (Doryctinae), Lissodoryctes (Doryctinae), Guaygata (Doryctinae), Tripteroides (Doryctinae), Donquickeia (Doryctinae), Waitaca (Doryctinae), Heerz (Doryctinae), Whartonius (Doryctinae), Heredius (Doryctinae), Coiba (Doryctinae), Hansonorum (Doryctinae), Subcurtisella (Doryctinae), Shawius (Doryctinae), Panama (Doryctinae), Platydoryctes (Doryctinae), Barbalhoa (Doryctinae), Aphelopsia (Doryctinae), Lianus (Doryctinae), Acanthorhogas (Doryctinae), Semirhytus (Doryctinae), Concurtisella (Doryctinae), Evaniodes (Doryctinae), Dapsilitas (Doryctinae), Iare (Doryctinae), Tripteria (Doryctinae), Johnsonius (Doryctinae), Harpoheterospilus (Doryctinae), Spathiospilus (Doryctinae), Jataiella (Doryctinae), Canchim (Doryctinae), Bohartiellus (Doryctinae), Caingangia (Doryctinae), Tarasco (Doryctinae), Masonius (Doryctinae), Whitfieldiellus (Doryctinae), Lamquetia (Doryctinae), Pedinotus (Doryctinae), Hybodoryctes (Doryctinae), Vanderentiellus (Doryctinae), Sericobracon (Doryctinae), Neostaphius (Doryctinae), Ondigus (Doryctinae), Trigonophasmus (Doryctinae), Osmophila (Doryctinae), Curtiselloides (Doryctinae), Oroceguera (Doryctinae), Ecphylopsis (Doryctinae), Termitospathius (Doryctinae), Australospathius (Doryctinae), Parana (Doryctinae), Ambispathius (Doryctinae), Antespathius (Doryctinae), Leptospathius (Doryctinae), Eopedinotus (Doryctinae), Arhaconotus (Doryctinae), Chelonodoryctes (Doryctinae), Mimipodoryctes (Doryctinae), Ryukyuspathius (Doryctinae), Ceylonspathius (Doryctinae), Halycaea (Doryctinae), Spathiostenus (Doryctinae), Paraspathius (Doryctinae), Synspilus (Doryctinae), Siragra (Doryctinae), Arkoola (Doryctinae), Pseudospathius (Doryctinae), Fijispathius (Doryctinae), Bracodoryctes (Doryctinae), Acanthodoryctes (Doryctinae), Pirramurra (Doryctinae), Ontsirospathius (Doryctinae), Neorhaconotus (Doryctinae), Dooliba (Doryctinae), Plyctes (Doryctinae), Liodoryctes (Doryctinae), Ceratodoryctes (Doryctinae), Fijibracon (Doryctinae), Pseudosyngaster (Doryctinae), Jarra (Doryctinae), Syngaster (Doryctinae), Antidoryctes (Doryctinae), Gurburra (Doryctinae), Echinodoryctes (Doryctinae), Doryctopsis (Doryctinae), Sisupala (Doryctinae), Preembobracon (Doryctinae), Doryctinus (Doryctinae), Neodoryctes (Doryctinae), Bolivar (Doryctinae), Tuberatra (Doryctinae), Venifurca (Doryctinae), Parsteres (Doryctinae), Sergey (Doryctinae), Asiaontsira (Doryctinae), Kauriphanes (Doryctinae), Papuaontsira (Doryctinae), Sycosoter (Doryctinae)
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith