Revision of Draft Key of the Doryctinae genera of the world for each geographic regions from Sun, 2014-11-02 12:56

This key is only a draft version provided to test the site. It has not been tested enough to confirm its quality. Moreover some errors could be present. So, if you wish to use it, please take in mind that all your identifications need to be seen with cautions and verified with additional sources!


The following genera have not been included because I lack data : Cecidospathius Kieffer & Jörgensen,1910; Celereon Say,1936; Doryctomorpha Asmead, 1900; Esterella Pagliano & Scaramozzino,1990; Metaspathius Brues, 1922; Sinaodoryctes Chen & Shi, 2004 (Key version 02/11/2014).

Draft - Key of Doryctinae of the world

Yves BRAET


Key 7. Confirmatory characters par regions geographiques

Characters: 137 in data, 99 included, 84 in key.
Items: 196 in data, 189 included, 379 in key.
Parameters: Rbase = 1.40 Abase = 2.00 Reuse = 1.01 Varywt = .30
Characters included: 2–44 46–57 59–89 92–101 118–119 137
Character reliabilities: 4,8.0 8,8.0 10–11,2.0 12,1.0 15,1.0 16,4.0 17,7.0 19,6.0 20,4.0 21–22,1.0 26,7.0 37–38,8.0 39,4.0 40–43,6.0 46,1.0 47,6.0 48,7.0 51,4.0 52–55,6.0 56,7.0 59,6.0 60–61,10.0 64,1.0 65,7.0 66–67,1.0 70,6.0 73,6.0 77,6.0 80,10.0 82,6.0 88,7.0 89,6.0 90–91,2.0 94,6.0 98,6.0 100–117,1.0 118–119,6.0 120–137,1.0

1(0).
  • Nearctic ... 2
  • Eastern Palearctic... 29
  • Western Palaearctic... 60
  • Afrotropical... 86
  • Neotropical... 128
  • Oriental... 241
  • Australasian... 288
  • Oceanic... 338

2(1).
  • Basoventral tooth of hind coxa absent ... 3
  • Basoventral tooth of hind coxa present (sometimes very small)... 11

3(2).
  • Apical lobe of scape absent; apical margin of scape and pedicel without dense cluster of setae; occipital carina present, at least laterally ... 4
  • Apical lobe of scape present; apical margin of scape and pedicel with dense cluster of setae; occipital carina absent... Liobracon Szépligeti, 1901

4(3).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... 5
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... 10
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long... Spathius Nees, 1918

5(4).
  • Propodeal bridge between abdominal and coxal foramina absent; hind tibia without comb of dense flat setae at inner apex ... 6
  • Propodeal bridge between abdominal and coxal foramina present; hind tibia with comb of dense flat setae at inner apex... Aivalykus Nixon, 1938

6(5).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; fore tibial spines usually few and forming a single row; scape shorter than first flagellomere ... 7
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; fore tibial spines more or less numerous and dispersed; scape equal to or longer than first flagellomere... Rhoptrocentrus Marshall, 1897

7(6).
  • Frons without lateral protuberances protruding to the surface of head ... 8
  • Frons with lateral protuberances protruding to the surface of head... Dendrosoter Wesmael, 1838 (including the subgenus)

8(7).
  • Hind coxa dorsally without teeth; basal area of second tergite enclosed by carinae or grooves absent; frons not excavated (flat or convex); raised ridge between antennae absent; third labial palp segment as long as or longer than second ... 9
  • Hind coxa with one to several teeth; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves); frons excavated (concave, sometimes weakly); raised ridge between antennae present; third labial palp segment distinctly shorter than second... Odontobracon Cameron, 1887

9(8).
  • Labial palpi four-segmented; maxillary palpi six-segmented ... Pambolidea Ashmead, 1900
  • Labial palpi three-segmented; maxillary palpi five-segmented... Ecphylus Foerster, 1862 (including the subgenus)

10(4).
  • First flagellar segment longer than second; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; occipital carina not meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Ptesimogaster Marsh, 1965
  • First flagellar segment equal to or shorter than second; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; occipital carina meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Pioscelus Muesebeck & Walkley, 1951

11(2).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... 12
  • Hind femora not swollen (at least 3x longer than wide)... 13

12(11).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); occipital carina not meeting hypostomal carina; propodeal carinated areas completely absent; propodeum with lateral tubercles (as high as large at base); first tergite from 1 times to 1.5 times longer than apical width ... Hemidoryctes Belokobylskij, 1993
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); occipital carina meeting hypostomal carina; propodeal carinated areas present at least basally; propodeum without lateral tubercles; first tergite less than 1 times longer than apical width... Stenocorse Marsh, 1968

13(11).
  • Wings present ... 14
  • Wings absent... Aptenobracon Marsh, 1965

14(13).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 15
  • Brachial (1st subdiscal) cell of fore wing open... 18

15(14).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 16
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)

16(15).
  • Recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); dorsope of first tergite absent or indistinct ... Spathius Nees, 1918
  • Recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); dorsope of first tergite present and more or less distinct... 17

17(16).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances ... Doryctes Haliday, 1836
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances... Ontsira Cameron, 1900

18(14).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 19
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 26

19(18).
  • First tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 20
  • First tergite apical width more than 1.6 times (even 2 times) basal width... 24

20(19).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts ... 21
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind wing male without a stigma-like enlargement... 23

21(20).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence ... 22
  • Metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body)... Glyptocolastes Ashmead, 1900

22(21).
  • Occipital carina meeting hypostomal carina; propodeum convex or abruptly declivous posteriorly; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused ... Heterospilus Halliday, 1836 (including the subgenus)
  • Occipital carina not meeting hypostomal carina; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated... Leluthia Cameron, 1887 (including the subgenus)

23(20).
  • First flagellar segment longer than second; second radiomedial vein (r-m) of fore wing present; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes smooth and shining ... Allorhogas Gahan, 1912
  • First flagellar segment equal to or shorter than second; second radiomedial vein (r-m) of fore wing absent; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); pronotum with convex lobe; mesoscutum surface of median and lateral lobes more or less sculptured... Monolexis Foerster, 1862

24(19).
  • Second radiomedial vein (r-m) of fore wing present; malar suture absent; notauli complete or at least partly present (smooth or sculptured); fore femur with more or less distinct anterodorsal protuberances; hind tibia with comb of dense flat setae at inner apex ... 25
  • Second radiomedial vein (r-m) of fore wing absent; malar suture present; notauli entirely absent; fore femur without anterodorsal protuberances; hind tibia without comb of dense flat setae at inner apex... Percnobraconoides Marsh, 1989

25(24).
  • Occipital carina meeting hypostomal carina; eyes setose; mesoscutum surface of median and lateral lobes more or less sculptured; scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron) ... Doryctinus Roman 1910
  • Occipital carina not meeting hypostomal carina; eyes glabrous; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron)... Parallorhogas Marsh, 1993

26(18).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M) ... 27
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M)... 28

27(26).
  • Occipital carina meeting hypostomal carina; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances ... Curtisella Spinola, 1853
  • Occipital carina not meeting hypostomal carina; ovipositor nodus double, with a second node weakly developed; hind wing male without a stigma-like enlargement; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances... Ptesimogaster Marsh, 1965

28(26).
  • Occipital carina meeting hypostomal carina; ocelar triangle equilateral ... Callihormius Ashmead, 1900b
  • Occipital carina not meeting hypostomal carina; ocelar triangle with base larger than its sides... Leluthia Cameron, 1887 (including the subgenus)

29(1).
  • Basoventral tooth of hind coxa absent ... 30
  • Basoventral tooth of hind coxa present (sometimes very small)... 42

30(29).
  • Propodeal bridge between abdominal and coxal foramina absent ... 31
  • Propodeal bridge between abdominal and coxal foramina present... Aivalykus Nixon, 1938

31(30).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 32
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 41

32(31).
  • Frons without lateral protuberances protruding to the surface of head ... 33
  • Frons with lateral protuberances protruding to the surface of head... Dendrosoter Wesmael, 1838 (including the subgenus)

33(32).
  • Fore tibial spines more or less numerous and dispersed ... 34
  • Fore tibial spines usually few and forming a single row... 35

34(33).
  • Occipital carina meeting hypostomal carina; hind femora not swollen (at least 3x longer than wide); ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully) ... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Occipital carina not meeting hypostomal carina; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); ovipositor nodus double, with a second node weakly developed; scutellum flat; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated)... Dendrosotinus Telenga, 1941

35(33).
  • Third tergite medially without any distinct transverse narrow depression/furrow ... 36
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... Pareucorystes Tobias, 1961

36(35).
  • First flagellar segment surface regularly smooth or finely sculptured ... 37
  • First flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view... Sonanus Belokobylskij & Konishi, 2001

37(36).
  • Basal area of second tergite enclosed by carinae or grooves absent ... 38
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... 39

38(37).
  • Propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width; eyes glabrous; scape shorter than first flagellomere; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more ... Ecphylus Foerster, 1862 (including the subgenus)
  • Propodeum with lateral tubercles (as high as large at base); first tergite less than 1 times longer than apical width; eyes setose; scape equal to or longer than first flagellomere; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined... Cryptontsira Belokobylskij, 2008

39(37).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; occipital carina present, at least laterally; hind coxa dorsally without teeth; third labial palp segment as long as or longer than second; raised ridge between antennae absent ... 40
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; occipital carina absent; hind coxa with one to several teeth; third labial palp segment distinctly shorter than second; raised ridge between antennae present... Zombrus Marshall, 1897

40(39).
  • First flagellar segment longer than second; second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally) ... Polystenus Förster, 1862
  • First flagellar segment equal to or shorter than second; second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent)... Eodendrus Belokobylskij, 1998

41(31).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; eyes setose; dorsope of first tergite present and more or less distinct ... Rhoptrocentrus Marshall, 1897
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; eyes glabrous; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

42(29).
  • Apical lobe of scape absent; hind wing without transverse vein (r) of radial cell ... 43
  • Apical lobe of scape present; hind wing with a transverse vein (r) of radial cell... Asiaheterospilus Belokobylskij & Konishi, 2001

43(42).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 44
  • Brachial (1st subdiscal) cell of fore wing open... 53

44(43).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Platyspathius Viereck, 1911
  • Hind femora not swollen (at least 3x longer than wide)... 45

45(44).
  • Fore wing of male without sclerotized enlargement, including veins 1-m and 1-SR+m ... 46
  • Fore wing of male with sclerotized enlargement... Neurocrassus Snoflak, 1945

46(45).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 47
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)

47(46).
  • Third tergite medially without any distinct transverse narrow depression/furrow; hind femora without anterodorsal protuberances; radial (marginal) cell of fore wing distally closed ... 48
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); hind femora with more or less distinct anterodorsal protuberances; radial (marginal) cell of fore wing open... Rhacontsira Belokobylskij, 1998

48(47).
  • Fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances ... 49
  • Fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances... 50

49(48).
  • Occipital carina meeting hypostomal carina; first tergite from 1.5 times to 2.5 times as long its apical width; mesoscutum surface of median and lateral lobes smooth and shining ... Hypodoryctes Kokujev, 1900
  • Occipital carina not meeting hypostomal carina; first tergite from 1 times to 1.5 times longer than apical width; mesoscutum surface of median and lateral lobes more or less sculptured... Ontsira Cameron, 1900

50(48).
  • Dorsope of first tergite present and more or less distinct ... 51
  • Dorsope of first tergite absent or indistinct... Spathius Nees, 1918

51(50).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; mesoscutum surface of median and lateral lobes more or less sculptured ... 52
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes smooth and shining... Doryctes Haliday, 1836

52(51).
  • First flagellar segment longer than second; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; propodeal carinated areas present at least basally; ovipositor nodus without visible node (but lower valves serrate); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron) ... Caenophanes Foerster, 1862 (including the subgenus)
  • First flagellar segment equal to or shorter than second; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; propodeal carinated areas completely absent; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron)... Eodendrus Belokobylskij, 1998

53(43).
  • Occipital carina meeting hypostomal carina ... 54
  • Occipital carina not meeting hypostomal carina... 57

54(53).
  • Second radiomedial vein (r-m) of fore wing present ... 55
  • Second radiomedial vein (r-m) of fore wing absent... 56

55(54).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scape shorter than first flagellomere; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; hind tibia without comb of dense flat setae at inner apex; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Heterospilus Halliday, 1836 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); scape equal to or longer than first flagellomere; hind wing male without a stigma-like enlargement; hind tibia with comb of dense flat setae at inner apex; first tergite apical width more than 1.6 times (even 2 times) basal width... Spathiomorpha Tobias, 1976

56(54).
  • First flagellar segment longer than second; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Hecabolus Curtis, 1837
  • First flagellar segment equal to or shorter than second; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind femora not swollen (at least 3x longer than wide)... Monolexis Foerster, 1862

57(53).
  • First flagellar segment longer than second; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); mid femora without anterodorsal protuberances; metapleuron glabrous or with a very sparse/rare discret pubescence ... 58
  • First flagellar segment equal to or shorter than second; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); ovipositor nodus without visible node (but lower valves serrate); mid femora with more or less distinct anterodorsal protuberances; metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body)... Hecabolomorpha Belokobylskij & Chen, 2006

58(57).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Hecabalodes Wilkinson, 1929
  • Hind femora not swollen (at least 3x longer than wide)... 59

59(58).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes more or less sculptured; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; fore femur without anterodorsal protuberances; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Leluthia Cameron, 1887 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes smooth and shining; hind wing male without a stigma-like enlargement; fore femur with more or less distinct anterodorsal protuberances; first tergite apical width more than 1.6 times (even 2 times) basal width... Parallorhogas Marsh, 1993

60(1).
  • Basoventral tooth of hind coxa absent ... 61
  • Basoventral tooth of hind coxa present (sometimes very small)... 70

61(60).
  • Frons without lateral protuberances protruding to the surface of head ... 62
  • Frons with lateral protuberances protruding to the surface of head... 69

62(61).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 63
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 68

63(62).
  • Third tergite medially without any distinct transverse narrow depression/furrow ... 64
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... Pareucorystes Tobias, 1961

64(63).
  • Fore tibial spines more or less numerous and dispersed; ovipositor nodus double, with a second node weakly developed ... Dendrosotinus Telenga, 1941
  • Fore tibial spines usually few and forming a single row; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak)... 65

65(64).
  • Basal area of second tergite enclosed by carinae or grooves absent ... 66
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... 67

66(65).
  • First flagellar segment longer than second; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); metanotum without any protuberances or spines; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; first tergite median carinae very short (less than or equal to the third of petiole length) ... Euscelinus Westwood, 1882
  • First flagellar segment equal to or shorter than second; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); metanotum with a short and acute erected spine medially; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; first tergite median carinae reaching, or nearly, the half length of the petiole... Ecphylus Foerster, 1862 (including the subgenus)

67(65).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; occipital carina present, at least laterally; hind coxa dorsally without teeth ... Polystenus Förster, 1862
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; occipital carina absent; hind coxa with one to several teeth... Zombrus Marshall, 1897

68(62).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; eyes setose; dorsope of first tergite present and more or less distinct ... Rhoptrocentrus Marshall, 1897
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; eyes glabrous; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

69(61).
  • Vein r of fore wing issuing clearly before middle of pterostigma; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); malar suture absent; scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated) ... Caenopachys Förster, 1862
  • Vein r of fore wing issuing submedially from pterostigma; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; malar suture present; scutellum flat; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully)... Dendrosoter Wesmael, 1838 (including the subgenus)

70(60).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 71
  • Brachial (1st subdiscal) cell of fore wing open... 79

71(70).
  • Second radiomedial vein (r-m) of fore wing present ... 72
  • Second radiomedial vein (r-m) of fore wing absent... 78

72(71).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); metanotum with a short and acute erected spine medially ... Platyspathius Viereck, 1911
  • Hind femora not swollen (at least 3x longer than wide); metanotum without any protuberances or spines... 73

73(72).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 74
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)

74(73).
  • Fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances ... 75
  • Fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances... 76

75(74).
  • Occipital carina meeting hypostomal carina; first tergite from 1.5 times to 2.5 times as long its apical width; mesoscutum surface of median and lateral lobes smooth and shining ... Hypodoryctes Kokujev, 1900
  • Occipital carina not meeting hypostomal carina; first tergite from 1 times to 1.5 times longer than apical width; mesoscutum surface of median and lateral lobes more or less sculptured... Ontsira Cameron, 1900

76(74).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; dorsope of first tergite present and more or less distinct ... 77
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

77(76).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused ... Caenophanes Foerster, 1862 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated... Doryctes Haliday, 1836

78(71).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); propodeal carinated areas completely absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind wing without transverse vein (r) of radial cell ... Mimodoryctes Belokobylskij, 2001
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); propodeal carinated areas present at least basally; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; hind wing with a transverse vein (r) of radial cell... Pseudorhaconotus van Achterberg & Shaw, 2010

79(70).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts ... 80
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind wing male without a stigma-like enlargement... 83

80(79).
  • Occipital carina meeting hypostomal carina ... 81
  • Occipital carina not meeting hypostomal carina... 82

81(80).
  • Second radiomedial vein (r-m) of fore wing present; hind femora not swollen (at least 3x longer than wide); scape shorter than first flagellomere; propodeum convex or abruptly declivous posteriorly; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused ... Heterospilus Halliday, 1836 (including the subgenus)
  • Second radiomedial vein (r-m) of fore wing absent; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); scape equal to or longer than first flagellomere; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated... Hecabolus Curtis, 1837

82(80).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Hecabalodes Wilkinson, 1929
  • Hind femora not swollen (at least 3x longer than wide)... Leluthia Cameron, 1887 (including the subgenus)

83(79).
  • First flagellar segment longer than second ... 84
  • First flagellar segment equal to or shorter than second... 85

84(83).
  • Occipital carina meeting hypostomal carina; hind tibia without comb of dense flat setae at inner apex; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; fore femur without anterodorsal protuberances; hind basitarsus up to 0.45 times as long as tarsi 2-5 combined ... Allorhogas Gahan, 1912
  • Occipital carina not meeting hypostomal carina; hind tibia with comb of dense flat setae at inner apex; first tergite apical width more than 1.6 times (even 2 times) basal width; fore femur with more or less distinct anterodorsal protuberances; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined... Parallorhogas Marsh, 1993

85(83).
  • Second radiomedial vein (r-m) of fore wing present; propodeum with lateral tubercles (as high as large at base); recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); eyes setose; pronotum dorsally without modifications ... Spathiomorpha Tobias, 1976
  • Second radiomedial vein (r-m) of fore wing absent; propodeum without lateral tubercles; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); eyes glabrous; pronotum with convex lobe... Monolexis Foerster, 1862

86(1).
  • Basoventral tooth of hind coxa absent ... 87
  • Basoventral tooth of hind coxa present (sometimes very small)... 111

87(86).
  • Propodeal bridge between abdominal and coxal foramina absent ... 88
  • Propodeal bridge between abdominal and coxal foramina present... 108

88(87).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 89
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 104

89(88).
  • Apical lobe of scape absent; scape less than two times longer than its maximum width ... 90
  • Apical lobe of scape present; scape two or more times longer than its maximum width... Rinamba Cameron,1912

90(89).
  • Frons without lateral protuberances protruding to the surface of head ... 91
  • Frons with lateral protuberances protruding to the surface of head... 102

91(90).
  • Fore tibial spines more or less numerous and dispersed ... 92
  • Fore tibial spines usually few and forming a single row... 94

92(91).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; first tergite from 1 times to 1.5 times longer than apical width; occipital carina present, at least laterally; metapleuron glabrous or with a very sparse/rare discret pubescence; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more ... 93
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; first tergite from 1.5 times to 2.5 times as long its apical width; occipital carina absent; metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body); hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined... Ivondrovia Shenefelt & Marsh, 1976

93(92).
  • Propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); third tergite medially without any distinct transverse narrow depression/furrow ... Dendrosotinus Telenga, 1941
  • Propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind femora not swollen (at least 3x longer than wide); third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... Gymnobracon Szépligeti, 1902

94(91).
  • Propodeum fully or partly (micro)sculptured; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances; dorsal spines of hind tibia absent ... 95
  • Propodeum almost entirely smooth (visible in the carinated areola); fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances; dorsal spines of hind tibia present... Doryctoproctus Belokobylskij, 2004

95(94).
  • Third tergite medially without any distinct transverse narrow depression/furrow; fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... 96
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface)... Odontodoryctes Granger, 1949

96(95).
  • Occipital carina present, at least laterally ... 97
  • Occipital carina absent... 101

97(96).
  • First flagellar segment longer than second ... 98
  • First flagellar segment equal to or shorter than second... 99

98(97).
  • Second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; propodeal carinated areas present at least basally; fore wing of male with sclerotized enlargement; hind wing male without a stigma-like enlargement ... Bulbonervus Shenefelt, 1969
  • Second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; propodeal carinated areas completely absent; fore wing of male without sclerotized enlargement, including veins 1-m and 1-SR+m; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts... Terate Nixon, 1943

99(97).
  • Hind coxa dorsally without teeth; first tergite without basolateral process (pointed wing-like, spine-like projections) near base ... 100
  • Hind coxa with one to several teeth; first tergite with basolateral process (pointed wing-like, spine-like projections) near base... Priosphys Enderlein, 1920

100(99).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); metanotum with a short and acute erected spine medially; hind tibia without comb of dense flat setae at inner apex; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Ecphylus Foerster, 1862 (including the subgenus)
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); metanotum without any protuberances or spines; hind tibia with comb of dense flat setae at inner apex; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; first tergite apical width more than 1.6 times (even 2 times) basal width... Eodendrus Belokobylskij, 1998

101(96).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); first tergite from 1.5 times to 2.5 times as long its apical width; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); second radiomedial vein (r-m) of fore wing with wide bulla ... Cryptodoryctes Belokobylskij & Quicke, 2000
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); first tergite from 1 times to 1.5 times longer than apical width; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); second radiomedial vein (r-m) of fore wing largely tubular... Zombrus Marshall, 1897

102(90).
  • First flagellar segment longer than second; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally) ... Spathioplites Fisher, 1962
  • First flagellar segment equal to or shorter than second; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent)... 103

103(102).
  • Vein r of fore wing issuing clearly before middle of pterostigma; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); malar suture absent ... Caenopachys Förster, 1862
  • Vein r of fore wing issuing submedially from pterostigma; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scutellum flat; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); malar suture present... Dendrosoter Wesmael, 1838 (including the subgenus)

104(88).
  • First flagellar segment longer than second; scutellum flat; notauli complete or at least partly present (smooth or sculptured) ... 105
  • First flagellar segment equal to or shorter than second; scutellum not flattened (e.g. swollen, conical, sharply raised); notauli entirely absent... Toka Nixon, 1943

105(104).
  • First flagellar segment surface regularly smooth or finely sculptured ... 106
  • First flagellar segment surface smooth dorsally, weakly curved and dorso-ventrally compressed... Paradoryctes Granger, 1949 (as synonym of Doryctes)
  • First flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view... Doryctophasmus Enderlein, 1912

106(105).
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; sternaulus (precoxal sulcus) absent; fore tibial spines absent ... Leptorhaconotus Granger, 1949
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; sternaulus (precoxal sulcus) present; fore tibial spines present... 107

107(106).
  • Propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); first tergite apical width more than 1.6 times (even 2 times) basal width; dorsope of first tergite absent or indistinct; basal area of second tergite enclosed by carinae or grooves absent ... Spathius Nees, 1918
  • Propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; dorsope of first tergite present and more or less distinct; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... Hemispathius Belokobylskij & Quicke, 2000

108(87).
  • Hind wing with the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing present; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind wing with the nervellus (cu-a) vein; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) ... 109
  • Hind wing without the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; hind wing without the nervellus (cu-a) vein; basal area of second tergite enclosed by carinae or grooves absent... Aivalykus Nixon, 1938

109(108).
  • Apical lobe of scape absent; first flagellar segment equal to or shorter than second; propodeal carinated areas completely absent; third tergite medially without any distinct transverse narrow depression/furrow; malar suture absent ... 110
  • Apical lobe of scape present; first flagellar segment longer than second; propodeal carinated areas present at least basally; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); malar suture present... Pseudodoryctes Szépligeti, 1915

110(109).
  • Frons without lateral protuberances protruding to the surface of head; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; basal area of second tergite separate from second suture; second radiomedial vein (r-m) of fore wing with wide bulla; first flagellar segment surface regularly smooth or finely sculptured ... Stephanospathius Belokobylskij, 1993
  • Frons with lateral protuberances protruding to the surface of head; acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; basal area of second tergite connected with second suture; second radiomedial vein (r-m) of fore wing largely tubular; first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view... Schlettereriella Szépligeti, 1904

111(86).
  • Hind wing with the recurrent vein (m-cu) ... 112
  • Hind wing without the recurrent vein (m-cu)... 127

112(111).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 113
  • Brachial (1st subdiscal) cell of fore wing open... 120

113(112).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 114
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... 119

114(113).
  • Recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent) ... 115
  • Recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally)... 117

115(114).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; dorsope of first tergite present and more or less distinct; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) ... Eodendrus Belokobylskij, 1998
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; dorsope of first tergite absent or indistinct; basal area of second tergite enclosed by carinae or grooves absent... 116

116(115).
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); metanotum with a short and acute erected spine medially; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances ... Platyspathius Viereck, 1911
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; hind femora not swollen (at least 3x longer than wide); metanotum without any protuberances or spines; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances... Spathius Nees, 1918

117(114).
  • Second tergite without apical lenticulate area ... 118
  • Second tergite with apical lenticulate area (often formed by the meeting of second suture and groove on third tergite when present)... Bathycentor Saussure, 1892

118(117).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances ... Doryctes Haliday, 1836
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances... Ontsira Cameron, 1900

119(113).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scape shorter than first flagellomere; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); first tergite median carinae reaching, or nearly, the half length of the petiole; fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scape equal to or longer than first flagellomere; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); first tergite median carinae more than the half length of petiole (sometimes reaching the apex of petiole); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface)... Ipodoryctes Granger, 1949

120(112).
  • Occipital carina meeting hypostomal carina ... 121
  • Occipital carina not meeting hypostomal carina... 125

121(120).
  • Second radiomedial vein (r-m) of fore wing present ... 122
  • Second radiomedial vein (r-m) of fore wing absent... Monolexis Foerster, 1862

122(121).
  • Scutellum flat; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 123
  • Scutellum not flattened (e.g. swollen, conical, sharply raised); first tergite apical width more than 1.6 times (even 2 times) basal width... 124

123(122).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; frons not excavated (flat or convex); propodeum convex or abruptly declivous posteriorly ... Heterospilus Halliday, 1836 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind wing male without a stigma-like enlargement; hind basitarsus up to 0.45 times as long as tarsi 2-5 combined; frons excavated (concave, sometimes weakly); propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... Allorhogas Gahan, 1912

124(122).
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); eyes setose; mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); scape shorter than first flagellomere ... Doryctinus Roman 1910
  • Ovipositor nodus modified apically (more or less widened); eyes glabrous; mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); scape equal to or longer than first flagellomere... Neoheterospilus Belokobylskij, 2006 (including the subgenera)

125(120).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; notauli complete or at least partly present (smooth or sculptured); scutellum flat; dorsope of first tergite present and more or less distinct; mid femora without anterodorsal protuberances ... 126
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; notauli entirely absent; scutellum not flattened (e.g. swollen, conical, sharply raised); dorsope of first tergite absent or indistinct; mid femora with more or less distinct anterodorsal protuberances... Afrospathius Belokobylskij & Quicke, 2000

126(125).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; second radiomedial vein (r-m) of fore wing present; propodeal carinated areas present at least basally; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind femora not swollen (at least 3x longer than wide) ... Parallorhogas Marsh, 1993
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; second radiomedial vein (r-m) of fore wing absent; propodeal carinated areas completely absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs)... Hecabalodes Wilkinson, 1929

127(111).
  • Second radiomedial vein (r-m) of fore wing present; hind femora not swollen (at least 3x longer than wide); first tergite from 1.5 times to 2.5 times as long its apical width; pronotum dorsally without modifications; scutellum flat ... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)
  • Second radiomedial vein (r-m) of fore wing absent; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); first tergite from 1 times to 1.5 times longer than apical width; pronotum with convex lobe; scutellum not flattened (e.g. swollen, conical, sharply raised)... Grangerdoryctes Belokobylskij, 2004

128(1).
  • Basoventral tooth of hind coxa absent ... 129
  • Basoventral tooth of hind coxa present (sometimes very small)... 173

129(128).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 130
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 164

130(129).
  • Apical lobe of scape absent ... 131
  • Apical lobe of scape present... 160

131(130).
  • Third tergite medially without any distinct transverse narrow depression/furrow ... 132
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... 157

132(131).
  • Sternaulus (precoxal sulcus) present ... 133
  • Sternaulus (precoxal sulcus) absent... 152

133(132).
  • Hind tibia without comb of dense flat setae at inner apex ... 134
  • Hind tibia with comb of dense flat setae at inner apex... 147

134(133).
  • First tergite less than 1 times longer than apical width ... 135
  • First tergite from 1 times to 1.5 times longer than apical width... 136
  • First tergite from 1.5 times to 2.5 times as long its apical width... 142

135(134).
  • First flagellar segment longer than second; brachial (1st subdiscal) cell of fore wing open; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; hind femora not swollen (at least 3x longer than wide) ... Monitoriella Hedqvist, 1963
  • First flagellar segment equal to or shorter than second; brachial (1st subdiscal) cell of fore wing distally closed; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs)... Cryptontsira Belokobylskij, 2008

136(134).
  • Scape equal to or longer than first flagellomere ... 137
  • Scape shorter than first flagellomere... 138

137(136).
  • Hind wing without transverse vein (r) of radial cell; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); raised ridge between antennae absent ... Holcobracon Cameron, 1905
  • Hind wing with a transverse vein (r) of radial cell; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); raised ridge between antennae present... Nervellius Roman, 1924

138(136).
  • Propodeum without lateral tubercles; hind coxa dorsally without teeth; basal area of second tergite enclosed by carinae or grooves absent ... 139
  • Propodeum with lateral tubercles (as high as large at base); hind coxa with one to several teeth; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... Odontobracon Cameron, 1887

139(138).
  • First tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 140
  • First tergite apical width more than 1.6 times (even 2 times) basal width... Glaucia Braga, Barbalho & Penteado-Dias, 2002

140(139).
  • First flagellar segment longer than second ... Mononeuron Fischer, 1981
  • First flagellar segment equal to or shorter than second... 141

141(140).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; metanotum with a short and acute erected spine medially; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; first tergite median carinae reaching, or nearly, the half length of the petiole; dorsope of first tergite present and more or less distinct ... Ecphylus Foerster, 1862 (including the subgenus)
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; metanotum without any protuberances or spines; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; first tergite median carinae more than the half length of petiole (sometimes reaching the apex of petiole); dorsope of first tergite absent or indistinct... Psenobolus Reinhard, 1885 (female and winged form)

142(134).
  • First flagellar segment longer than second; brachial (1st subdiscal) cell of fore wing distally closed; pronotum with convex lobe; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); fore femur with more or less distinct anterodorsal protuberances ... 143
  • First flagellar segment equal to or shorter than second; brachial (1st subdiscal) cell of fore wing open; pronotum dorsally without modifications; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); fore femur without anterodorsal protuberances... 144

143(142).
  • Hind wing with the recurrent vein (m-cu); propodeal bridge between abdominal and coxal foramina absent; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeum fully or partly (micro)sculptured; propodeal carinated areas present at least basally ... Ficobolus Martínez, Belokobylskij et Zaldívar-Riverón, 2014
  • Hind wing without the recurrent vein (m-cu); propodeal bridge between abdominal and coxal foramina present; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; propodeum almost entirely smooth (visible in the carinated areola); propodeal carinated areas completely absent... Monarea Szépligeti, 1904

144(142).
  • Second radiomedial vein (r-m) of fore wing present; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); fore tibial spines usually few and forming a single row; hind wing with the nervellus (cu-a) vein; medial (basal) cell of hind wing closed ... 145
  • Second radiomedial vein (r-m) of fore wing absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines more or less numerous and dispersed; hind wing without the nervellus (cu-a) vein; medial (basal) cell of hind wing widely open antero-distally (vein SC+R absent)... 146

145(144).
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); hind femora not swollen (at least 3x longer than wide); mesoscutum surface of median and lateral lobes more or less sculptured; second radiomedial vein (r-m) of fore wing largely tubular ... Pioscelus Muesebeck & Walkley, 1951
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); mesoscutum surface of median and lateral lobes smooth and shining; second radiomedial vein (r-m) of fore wing with wide bulla... Psenobolus Reinhard, 1885 (female and winged form)

146(144).
  • Propodeal carinated areas present at least basally; nervellus (cu-a) of fore wing antefurcal; occipital carina meeting hypostomal carina; notauli entirely absent; scutellum not flattened (e.g. swollen, conical, sharply raised) ... Percnobracon Kieffer & Jörgensen, 1910
  • Propodeal carinated areas completely absent; nervellus (cu-a) of fore wing postfurcal or interstitial; occipital carina not meeting hypostomal carina; notauli complete or at least partly present (smooth or sculptured); scutellum flat... Micrommatus Marsh, 1993

147(133).
  • Hind wing with the recurrent vein (m-cu); occipital carina meeting hypostomal carina ... 148
  • Hind wing without the recurrent vein (m-cu); occipital carina not meeting hypostomal carina... 151

148(147).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing ... 149
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing... Bolivar Zaldívar-Riverón & Rodríguez-Jiménez, 2013

149(148).
  • First flagellar segment longer than second; second radiomedial vein (r-m) of fore wing present; eyes glabrous; hind wing with the nervellus (cu-a) vein ... 150
  • First flagellar segment equal to or shorter than second; second radiomedial vein (r-m) of fore wing absent; eyes setose; hind wing without the nervellus (cu-a) vein... Fritziella Marsh, 1993

150(149).
  • Ovipositor nodus without visible node (but lower valves serrate); metanotum without any protuberances or spines; metapleuron glabrous or with a very sparse/rare discret pubescence; hind basitarsus up to 0.45 times as long as tarsi 2-5 combined; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Heterospathius Barbalho & Penteado-Dias, 1999
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); metanotum with a short and acute erected spine medially; metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body); hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; first tergite apical width more than 1.6 times (even 2 times) basal width... Notiospathius Matthews & Marsh, 1973

151(147).
  • Propodeal bridge between abdominal and coxal foramina absent; brachial (1st subdiscal) cell of fore wing open; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeum with lateral tubercles (as high as large at base); first tergite from 1.5 times to 2.5 times as long its apical width ... Leptodoryctes Barbalho & Penteado-Dias, 1999
  • Propodeal bridge between abdominal and coxal foramina present; brachial (1st subdiscal) cell of fore wing distally closed; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width... Aivalykus Nixon, 1938

152(132).
  • First flagellar segment longer than second; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 153
  • First flagellar segment equal to or shorter than second; propodeum convex or abruptly declivous posteriorly... Termitobracon Brues, 1923

153(152).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 154
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; first tergite apical width more than 1.6 times (even 2 times) basal width... 156

154(153).
  • Hind wing with the recurrent vein (m-cu); first tergite from 1 times to 1.5 times longer than apical width; occipital carina present, at least laterally; notauli complete or at least partly present (smooth or sculptured); epicnemial carina present ... 155
  • Hind wing without the recurrent vein (m-cu); first tergite less than 1 times longer than apical width; occipital carina absent; notauli entirely absent; epicnemial carina absent... Ypsistocerus Cushman, 1923

155(154).
  • Propodeum fully or partly (micro)sculptured; propodeal carinated areas completely absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scape shorter than first flagellomere; hind wing without the nervellus (cu-a) vein ... Labania Hedqvist, 1963
  • Propodeum almost entirely smooth (visible in the carinated areola); propodeal carinated areas present at least basally; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scape equal to or longer than first flagellomere; hind wing with the nervellus (cu-a) vein... Histeromeroides Marsh, 1993

156(153).
  • Second radiomedial vein (r-m) of fore wing present; occipital carina not meeting hypostomal carina; hind femora not swollen (at least 3x longer than wide); first tergite from 1 times to 1.5 times longer than apical width; ovipositor nodus double, with a second node weakly developed ... Embobracon van Achterberg, 1995
  • Second radiomedial vein (r-m) of fore wing absent; occipital carina meeting hypostomal carina; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); first tergite from 1.5 times to 2.5 times as long its apical width; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak)... Verae Marsh, 1993

157(131).
  • Brachial (1st subdiscal) cell of fore wing distally closed; occipital carina not meeting hypostomal carina ... 158
  • Brachial (1st subdiscal) cell of fore wing open; occipital carina meeting hypostomal carina... 159

158(157).
  • Propodeal bridge between abdominal and coxal foramina absent; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); hind wing without transverse vein (r) of radial cell; fore tibial spines more or less numerous and dispersed; pronotum dorsally without modifications ... Gymnobracon Szépligeti, 1902
  • Propodeal bridge between abdominal and coxal foramina present; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); hind wing with a transverse vein (r) of radial cell; fore tibial spines usually few and forming a single row; pronotum with convex lobe... Megaloproctus Schulz, 1906

159(157).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines more or less numerous and dispersed; mesoscutum surface of median and lateral lobes more or less sculptured; hind tibia with comb of dense flat setae at inner apex; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Amazondoryctes Barbalho & Penteado-Dias, 1999
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); fore tibial spines usually few and forming a single row; mesoscutum surface of median and lateral lobes smooth and shining; hind tibia without comb of dense flat setae at inner apex; first tergite apical width more than 1.6 times (even 2 times) basal width... Pannuceus Marsh, 2002

160(130).
  • Hind wing with the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing present; hind wing with the nervellus (cu-a) vein; first tergite with dorsal carina (one or two, often mediolongitudinally or submediolongitudinally)(if the tergite is fully longitudinally striate, this(these) carina(e) could be hardly distinguished from the other and are coded as present); second metasomal suture present ... 161
  • Hind wing without the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing absent; hind wing without the nervellus (cu-a) vein; first tergite without dorsal carinae; second metasomal suture largely or entirely absent... Achterbergia Marsh, 1993

161(160).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 162
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes more or less sculptured; scutellum not flattened (e.g. swollen, conical, sharply raised); propodeum convex or abruptly declivous posteriorly... 163

162(161).
  • First tergite less than 1 times longer than apical width ... Liobracon Szépligeti, 1901
  • First tergite from 1 times to 1.5 times longer than apical width... Rinamba Cameron,1912
  • First tergite from 1.5 times to 2.5 times as long its apical width... Pannuceus Marsh, 2002

163(161).
  • Frons without lateral protuberances protruding to the surface of head; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); first tergite from 1.5 times to 2.5 times as long its apical width; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak) ... Plesiopsenobolus Belokobylskij, Martínez et Zaldívar-Riverón, 2014
  • Frons with lateral protuberances protruding to the surface of head; acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); first tergite from 1 times to 1.5 times longer than apical width; ovipositor nodus without visible node (but lower valves serrate)... Sabinita Belokobylskij, Zaldívar-Riverón & Martínez, 2014

164(129).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... 165
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... 166
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long... 171

165(164).
  • First tergite from 1 times to 1.5 times longer than apical width ... Binarea Brullé, 1846
  • First tergite from 1.5 times to 2.5 times as long its apical width... Dicarinoryctes Braet & van Achterberg, 2001
  • First tergite more 2.5 times its apical width... Rimacollus Marsh, 2002

166(164).
  • First flagellar segment longer than second; second metasomal suture present ... 167
  • First flagellar segment equal to or shorter than second; second metasomal suture largely or entirely absent... 169

167(166).
  • Propodeal carinated areas present at least basally; propodeum with lateral long spines (higher than its base); eyes setose; dorsope of first tergite absent or indistinct; mesoscutum surface of median and lateral lobes smooth and shining ... Doryctopambolus Nunes & Zaldívar-Riverón, 2012
  • Propodeal carinated areas completely absent; propodeum without lateral tubercles; eyes glabrous; dorsope of first tergite present and more or less distinct; mesoscutum surface of median and lateral lobes more or less sculptured... 168

168(167).
  • Occipital carina meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); fore tibial spines more or less numerous and dispersed; pronotum with convex lobe; first tergite without basolateral process (pointed wing-like, spine-like projections) near base ... Janzenia Marsh, 1993
  • Occipital carina not meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); fore tibial spines usually few and forming a single row; pronotum dorsally without modifications; first tergite with basolateral process (pointed wing-like, spine-like projections) near base... Ptesimogaster Marsh, 1965

169(166).
  • Propodeum fully or partly (micro)sculptured; hind femora not swollen (at least 3x longer than wide); occipital carina present, at least laterally; scutellum not flattened (e.g. swollen, conical, sharply raised); epicnemial carina present ... 170
  • Propodeum almost entirely smooth (visible in the carinated areola); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); occipital carina absent; scutellum flat; epicnemial carina absent... Psenobolus Reinhard, 1885 (male and brachypterous form)

170(169).
  • Fore tibial spines more or less numerous and dispersed; first tergite from 1.5 times to 2.5 times as long its apical width; propodeum convex or abruptly declivous posteriorly ... Oroceguera Seltmann & Sharkey, 2007
  • Fore tibial spines usually few and forming a single row; first tergite from 1 times to 1.5 times longer than apical width; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... Ecphylopsis Ashmead, 1900

171(164).
  • Hind wing with the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing present; notauli complete or at least partly present (smooth or sculptured) ... 172
  • Hind wing without the recurrent vein (m-cu); second radiomedial vein (r-m) of fore wing absent; notauli entirely absent... Angelica Marsh, 2002

172(171).
  • First flagellar segment longer than second; brachial (1st subdiscal) cell of fore wing distally closed; propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); third tergite medially without any distinct transverse narrow depression/furrow ... Spathius Nees, 1918
  • First flagellar segment equal to or shorter than second; brachial (1st subdiscal) cell of fore wing open; propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... Sharkeyella Marsh, 1993

173(128).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 174
  • Brachial (1st subdiscal) cell of fore wing open... 190

174(173).
  • Hind wing with the recurrent vein (m-cu) ... 175
  • Hind wing without the recurrent vein (m-cu)... 189

175(174).
  • Nervellus (cu-a) of fore wing postfurcal or interstitial ... 176
  • Nervellus (cu-a) of fore wing antefurcal... Hybodoryctes Szépligeti, 1906

176(175).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... 177
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A)... 178
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 179

177(176).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; dorsope of first tergite present and more or less distinct; fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally ... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; dorsope of first tergite absent or indistinct; fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth... Spathius Nees, 1918

178(176).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; dorsope of first tergite present and more or less distinct ... Guaygata Marsh, 1993
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); first tergite apical width more than 1.6 times (even 2 times) basal width; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

179(176).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing ... 180
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing... 187

180(179).
  • Occipital carina meeting hypostomal carina ... 181
  • Occipital carina not meeting hypostomal carina... 184

181(180).
  • First tergite from 1 times to 1.5 times longer than apical width ... 182
  • First tergite from 1.5 times to 2.5 times as long its apical width... 183
  • First tergite more 2.5 times its apical width... Lianus Gomes & Penteado-Dias, 2006

182(181).
  • Propodeum fully or partly (micro)sculptured; mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa); hind tibia with comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves); second radiomedial vein (r-m) of fore wing largely tubular ... Lamquetia Braet, Barbalhoa & van Achterberg, 2003
  • Propodeum almost entirely smooth (visible in the carinated areola); mesoscutum surface of median and lateral lobes smooth and shining; hind tibia without comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves absent; second radiomedial vein (r-m) of fore wing with wide bulla... Lissodoryctes Marsh, 2002

183(181).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); propodeal carinated areas completely absent; fore tibial spines usually few and forming a single row; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); eyes glabrous ... Acanthorhogas Szepligeti, 1906
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); propodeal carinated areas present at least basally; fore tibial spines more or less numerous and dispersed; third tergite medially without any distinct transverse narrow depression/furrow; eyes setose... Hypodoryctes Kokujev, 1900

184(180).
  • First flagellar segment longer than second; dorsope of first tergite present and more or less distinct ... 185
  • First flagellar segment equal to or shorter than second; dorsope of first tergite absent or indistinct... Trigonophasmus Enderlein, 1912

185(184).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) ... 186
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; basal area of second tergite enclosed by carinae or grooves absent... Cyphodoryctes Marsh, 1997

186(185).
  • Mesoscutum surface of median and lateral lobes smooth and shining; second radiomedial vein (r-m) of fore wing largely tubular; scape shorter than first flagellomere; hind tibia without comb of dense flat setae at inner apex; first tergite without basolateral process (pointed wing-like, spine-like projections) near base ... Pedinotus Szépligeti, 1902 (including the subgenus Eopedinotus)
  • Mesoscutum surface of median and lateral lobes more or less sculptured; second radiomedial vein (r-m) of fore wing with wide bulla; scape equal to or longer than first flagellomere; hind tibia with comb of dense flat setae at inner apex; first tergite with basolateral process (pointed wing-like, spine-like projections) near base... Osmophila Szépligeti, 1902

187(179).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); scape shorter than first flagellomere; hind wing male without a stigma-like enlargement; third labial palp segment as long as or longer than second ... 188
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); scape equal to or longer than first flagellomere; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; third labial palp segment distinctly shorter than second... Araucania Marsh, 1993

188(187).
  • Fore tibial spines more or less numerous and dispersed; second radiomedial vein (r-m) of fore wing with wide bulla; hind tibia with comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves absent ... Tripteria Enderlein, 1912
  • Fore tibial spines usually few and forming a single row; second radiomedial vein (r-m) of fore wing largely tubular; hind tibia without comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... Pedinotus Szépligeti, 1902 (including the subgenus Eopedinotus)

189(174).
  • Second radiomedial vein (r-m) of fore wing present; propodeum without median process; hind wing with the nervellus (cu-a) vein; second metasomal suture present; fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally ... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)
  • Second radiomedial vein (r-m) of fore wing absent; propodeum with a medio-longitudinal raised lamella; hind wing without the nervellus (cu-a) vein; second metasomal suture largely or entirely absent; fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth... Bohartiellus Marsh, 1983

190(173).
  • Hind wing with the recurrent vein (m-cu) ... 191
  • Hind wing without the recurrent vein (m-cu)... 239

191(190).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 192
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 233

192(191).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing ... 193
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing... 226

193(192).
  • Propodeum without median process ... 194
  • Propodeum with a median raised tooth or tubercle... Coiba Marsh, 1993

194(193).
  • Second radiomedial vein (r-m) of fore wing present ... 195
  • Second radiomedial vein (r-m) of fore wing absent... 220

195(194).
  • Mid femora with more or less distinct anterodorsal protuberances; hind femora with more or less distinct anterodorsal protuberances ... 196
  • Mid femora without anterodorsal protuberances; hind femora without anterodorsal protuberances... 200

196(195).
  • Propodeal bridge between abdominal and coxal foramina absent; first flagellar segment longer than second ... 197
  • Propodeal bridge between abdominal and coxal foramina present; first flagellar segment equal to or shorter than second... Neostaphius Braet, Barbalhoa & van Achterberg, 2003

197(196).
  • Propodeum fully or partly (micro)sculptured; mesoscutum surface of median and lateral lobes more or less sculptured ... 198
  • Propodeum almost entirely smooth (visible in the carinated areola); mesoscutum surface of median and lateral lobes smooth and shining... Semirhytus Szépligeti, 1902

198(197).
  • Ovipositor nodus single ... Barbalhoa Marsh, 2002
  • Ovipositor nodus double, with a second node weakly developed... 199
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak)... Doryctinus Roman 1910

199(198).
  • Fore tibial spines more or less numerous and dispersed; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body); scape shorter than first flagellomere ... Concurtisella Roman, 1924
  • Fore tibial spines usually few and forming a single row; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron glabrous or with a very sparse/rare discret pubescence; scape equal to or longer than first flagellomere... Ondigus Braet, Barbalhoa & van Achterberg, 2003

200(195).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; dorsope of first tergite present and more or less distinct ... 201
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; dorsope of first tergite absent or indistinct... 218

201(200).
  • Occipital carina meeting hypostomal carina ... 202
  • Occipital carina not meeting hypostomal carina... 213

202(201).
  • First tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 203
  • First tergite apical width more than 1.6 times (even 2 times) basal width... 212

203(202).
  • Shape of second metasomal suture straight or evenly curved ... 204
  • Shape of second metasomal suture with more or less distinct lateral angulations... 209

204(203).
  • Fore tibial spines absent; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) ... Sericobracon Shaw & Edgerly, 1985
  • Fore tibial spines present; basal area of second tergite enclosed by carinae or grooves absent... 205

205(204).
  • Propodeum fully or partly (micro)sculptured ... 206
  • Propodeum almost entirely smooth (visible in the carinated areola)... Waitaca Marsh, 1993

206(205).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A) ... 207
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 208

207(206).
  • Propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded); first radiomedial vein (2-SR) of fore wing present, leading the first and second submarginal cells of fore wing separated; frons excavated (concave, sometimes weakly) ... Heredius Marsh, 2002
  • Propodeum convex or abruptly declivous posteriorly; first radiomedial vein (2-SR) of fore wing absent or strongly unsclerotized for the most part, then first and second submarginal cells of fore wing fused; frons not excavated (flat or convex)... Heterospilus Halliday, 1836 (including the subgenus)

208(206).
  • First tergite less than 1 times longer than apical width; scape equal to or longer than first flagellomere; frons not excavated (flat or convex); vertex smooth ... Heerz Marsh, 1993
  • First tergite from 1 times to 1.5 times longer than apical width; scape shorter than first flagellomere; frons excavated (concave, sometimes weakly); vertex granulate, rugulose-granulate, acinose or coriaceous... Allorhogas Gahan, 1912

209(203).
  • First flagellar segment longer than second; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally) ... 210
  • First flagellar segment equal to or shorter than second; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent)... Donquickeia Marsh, 1997

210(209).
  • Hind tibia without comb of dense flat setae at inner apex; scape two or more times longer than its maximum width ... Whartonius Marsh, 1993
  • Hind tibia with comb of dense flat setae at inner apex; scape less than two times longer than its maximum width... 211

211(210).
  • Propodeal carinated areas present at least basally; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); notauli entirely absent ... Stenocorse Marsh, 1968
  • Propodeal carinated areas completely absent; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind femora not swollen (at least 3x longer than wide); ovipositor nodus double, with a second node weakly developed; notauli complete or at least partly present (smooth or sculptured)... Glyptocolastes Ashmead, 1900

212(202).
  • First flagellar segment longer than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; scutellum not flattened (e.g. swollen, conical, sharply raised) ... Neoheterospilus Belokobylskij, 2006 (including the subgenera)
  • First flagellar segment equal to or shorter than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; scutellum flat... Vanderentiellus Marsh, 2002

213(201).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Hemidoryctes Belokobylskij, 1993
  • Hind femora not swollen (at least 3x longer than wide)... 214

214(213).
  • Mesoscutum surface of median and lateral lobes smooth and shining ... 215
  • Mesoscutum surface of median and lateral lobes more or less sculptured... 216

215(214).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); fore femur with more or less distinct anterodorsal protuberances; first tergite apical width more than 1.6 times (even 2 times) basal width; first tergite median carinae very short (less than or equal to the third of petiole length) ... Parallorhogas Marsh, 1993
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); fore femur without anterodorsal protuberances; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; first tergite median carinae reaching, or nearly, the half length of the petiole... Dapsilitas Braet, Barbalhoa & van Achterberg, 2003

216(214).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M) ... Tripteroides Marsh, 2002
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M)... 217

217(216).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; fore tibial spines present ... Leluthia Cameron, 1887 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); hind wing male without a stigma-like enlargement; fore tibial spines absent... Sericobracon Shaw & Edgerly, 1985

218(200).
  • Propodeal bridge between abdominal and coxal foramina absent; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 219
  • Propodeal bridge between abdominal and coxal foramina present; propodeum convex or abruptly declivous posteriorly... Evaniodes Szépligeti, 1901

219(218).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); propodeal carinated areas completely absent; hind wing male without a stigma-like enlargement; hind tibia with comb of dense flat setae at inner apex; first tergite apical width more than 1.6 times (even 2 times) basal width ... Notiospathius Matthews & Marsh, 1973
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); propodeal carinated areas present at least basally; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; hind tibia without comb of dense flat setae at inner apex; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width... Spathiospilus Marsh, 1999

220(194).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... 221
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A)... 223
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 224

221(220).
  • First tergite from 1 times to 1.5 times longer than apical width; hind tibia without comb of dense flat setae at inner apex; first tergite apical width more than 1.6 times (even 2 times) basal width ... 222
  • First tergite more 2.5 times its apical width; hind tibia with comb of dense flat setae at inner apex; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width... Tarasco Marsh, 1993

222(221).
  • First flagellar segment longer than second; fore tibial spines more or less numerous and dispersed; notauli entirely absent; scutellum flat; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron) ... Percnobraconoides Marsh, 1989
  • First flagellar segment equal to or shorter than second; fore tibial spines usually few and forming a single row; notauli complete or at least partly present (smooth or sculptured); scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron)... Masonius Marsh, 1993

223(220).
  • Occipital carina meeting hypostomal carina; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); scape equal to or longer than first flagellomere ... Hecabolus Curtis, 1837
  • Occipital carina not meeting hypostomal carina; hind femora not swollen (at least 3x longer than wide); scape shorter than first flagellomere... Leluthia Cameron, 1887 (including the subgenus)

224(220).
  • Third tergite medially without any distinct transverse narrow depression/furrow; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); scape less than two times longer than its maximum width; frons not excavated (flat or convex) ... 225
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); scape two or more times longer than its maximum width; frons excavated (concave, sometimes weakly)... Caingangia Marsh, 1993

225(224).
  • Pronotum with convex lobe; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); hind tibia with comb of dense flat setae at inner apex; shape of second metasomal suture with more or less distinct lateral angulations; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... Monolexis Foerster, 1862
  • Pronotum dorsally without modifications; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); hind tibia without comb of dense flat setae at inner apex; shape of second metasomal suture straight or evenly curved; propodeum convex or abruptly declivous posteriorly... Canchim Barbalho & Penteado-Dias, 1999

226(192).
  • Second radiomedial vein (r-m) of fore wing present ... 227
  • Second radiomedial vein (r-m) of fore wing absent... Lissopius Marsh, 2002

227(226).
  • Apical lobe of scape absent; propodeal carinated areas present at least basally; scape less than two times longer than its maximum width ... 228
  • Apical lobe of scape present; propodeal carinated areas completely absent; scape two or more times longer than its maximum width... Whitfieldiellus Marsh, 1997

228(227).
  • Sternaulus (precoxal sulcus) present; second radiomedial vein (r-m) of fore wing with wide bulla ... 229
  • Sternaulus (precoxal sulcus) absent; second radiomedial vein (r-m) of fore wing largely tubular... Jataiella Barbalho & Penteado-Dias, 1999

229(228).
  • Mid femora with more or less distinct anterodorsal protuberances ... Johnsonius Marsh, 1993
  • Mid femora without anterodorsal protuberances... 230

230(229).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A) ... Glaucia Braga, Barbalho & Penteado-Dias, 2002
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 231

231(230).
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); pronotum dorsally without modifications; scutellum flat; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); scape shorter than first flagellomere ... 232
  • Ovipositor nodus modified apically (more or less widened); pronotum with convex lobe; scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); scape equal to or longer than first flagellomere... Neoheterospilus Belokobylskij, 2006 (including the subgenera)

232(231).
  • Occipital carina meeting hypostomal carina; fore femur without anterodorsal protuberances; hind tibia without comb of dense flat setae at inner apex; hind basitarsus up to 0.45 times as long as tarsi 2-5 combined; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Allorhogas Gahan, 1912
  • Occipital carina not meeting hypostomal carina; fore femur with more or less distinct anterodorsal protuberances; hind tibia with comb of dense flat setae at inner apex; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; first tergite apical width more than 1.6 times (even 2 times) basal width... Parallorhogas Marsh, 1993

233(191).
  • Occipital carina meeting hypostomal carina ... 234
  • Occipital carina not meeting hypostomal carina... 236

234(233).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); pronotum dorsally without modifications; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances; hind tibia with comb of dense flat setae at inner apex ... Curtisella Spinola, 1853
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); pronotum with convex lobe; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances; hind tibia without comb of dense flat setae at inner apex... 235

235(234).
  • Scape equal to or longer than first flagellomere; scape two or more times longer than its maximum width; second tergite without apical lenticulate area; fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface); vertex striate ... Shawius Marsh, 1993
  • Scape shorter than first flagellomere; scape less than two times longer than its maximum width; second tergite with apical lenticulate area (often formed by the meeting of second suture and groove on third tergite when present); fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly; vertex not striate... Callihormius Ashmead, 1900b

236(233).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... 237
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... 238

237(236).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); first and second tergites fused; ovipositor nodus double, with a second node weakly developed; scutellum not flattened (e.g. swollen, conical, sharply raised); fore femur with more or less distinct anterodorsal protuberances ... Iare Barbalho & Penteado-Dias, 2002
  • Hind femora not swollen (at least 3x longer than wide); first and second tergites not fused; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); scutellum flat; fore femur without anterodorsal protuberances... Leluthia Cameron, 1887 (including the subgenus)

238(236).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scutellum concave ... Callihormius (Dmitriohormius) Belokobylskij, Zaldívar-Riverón & Coronado-Blanco, 2014
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scutellum not flattened (e.g. swollen, conical, sharply raised)... Aphelopsia Marsh, 1993
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); scutellum flat... Ptesimogaster Marsh, 1965

239(190).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; notauli entirely absent ... Platydoryctes Barbalhoa & Penteado-Dias, 2000
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); notauli complete or at least partly present (smooth or sculptured)... 240

240(239).
  • Ovipositor nodus single ... Curtiselloides Marsh, 2002
  • Ovipositor nodus double, with a second node weakly developed... Ptesimogaster Marsh, 1965
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak)... Curtisella Spinola, 1853

241(1).
  • Apical lobe of scape absent ... 242
  • Apical lobe of scape present... 285

242(241).
  • Basoventral tooth of hind coxa absent ... 243
  • Basoventral tooth of hind coxa present (sometimes very small)... 260

243(242).
  • Propodeal bridge between abdominal and coxal foramina absent ... 244
  • Propodeal bridge between abdominal and coxal foramina present... 259

244(243).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 245
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 255

245(244).
  • Propodeum fully or partly (micro)sculptured; dorsope of first tergite present and more or less distinct; subapical teeth on fore and middle femora ventrally absent ... 246
  • Propodeum almost entirely smooth (visible in the carinated areola); dorsope of first tergite absent or indistinct; subapical teeth on fore and middle femora ventrally present... Termitospathius Belokobylskij, 2002

246(245).
  • Frons without lateral protuberances protruding to the surface of head ... 247
  • Frons with lateral protuberances protruding to the surface of head... Dendrosoter Wesmael, 1838 (including the subgenus)

247(246).
  • Fore tibial spines more or less numerous and dispersed ... Dendrosotinus Telenga, 1941
  • Fore tibial spines usually few and forming a single row... 248

248(247).
  • Basal area of second tergite enclosed by carinae or grooves absent ... 249
  • Basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... 252

249(248).
  • First flagellar segment longer than second; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron) ... 250
  • First flagellar segment equal to or shorter than second; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron)... 251

250(249).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; propodeal carinated areas completely absent; propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width; first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view ... Euscelinus Westwood, 1882
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeal carinated areas present at least basally; propodeum with lateral tubercles (as high as large at base); first tergite from 1.5 times to 2.5 times as long its apical width; first flagellar segment surface regularly smooth or finely sculptured... Spathius (Ambispathius) Belokobylskij, 1995

251(249).
  • Propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width; eyes glabrous; scape shorter than first flagellomere; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more ... Ecphylus Foerster, 1862 (including the subgenus)
  • Propodeum with lateral tubercles (as high as large at base); first tergite less than 1 times longer than apical width; eyes setose; scape equal to or longer than first flagellomere; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined... Cryptontsira Belokobylskij, 2008

252(248).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; third labial palp segment as long as or longer than second ... 253
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; third labial palp segment distinctly shorter than second... 254

253(252).
  • First flagellar segment longer than second; second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally) ... Polystenus Förster, 1862
  • First flagellar segment equal to or shorter than second; second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent)... Eodendrus Belokobylskij, 1998

254(252).
  • Occipital carina present, at least laterally; second radiomedial vein (r-m) of fore wing with wide bulla; hind coxa dorsally without teeth; frons not excavated (flat or convex); raised ridge between antennae absent ... Holcobracon Cameron, 1905
  • Occipital carina absent; second radiomedial vein (r-m) of fore wing largely tubular; hind coxa with one to several teeth; frons excavated (concave, sometimes weakly); raised ridge between antennae present... Zombrus Marshall, 1897

255(244).
  • Hind wing with the recurrent vein (m-cu); first flagellar segment longer than second ... 256
  • Hind wing without the recurrent vein (m-cu); first flagellar segment equal to or shorter than second... 258

256(255).
  • Nervellus (cu-a) of fore wing postfurcal or interstitial; scutellum flat; scape less than two times longer than its maximum width; first tergite apical width more than 1.6 times (even 2 times) basal width; dorsope of first tergite absent or indistinct ... 257
  • Nervellus (cu-a) of fore wing antefurcal; scutellum not flattened (e.g. swollen, conical, sharply raised); scape two or more times longer than its maximum width; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; dorsope of first tergite present and more or less distinct... Spathius (Antespathius) Belokobylskij, 1995

257(256).
  • Propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); first flagellar segment surface regularly smooth or finely sculptured; hind wing male without a stigma-like enlargement; dorsal spines of hind tibia absent ... Spathius Nees, 1918
  • Propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; dorsal spines of hind tibia present... Doryctophasmus Enderlein, 1912

258(255).
  • Second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); nervellus (cu-a) of fore wing antefurcal; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... Parana Nixon, 1943
  • Second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); nervellus (cu-a) of fore wing postfurcal or interstitial; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... Spathiostenus Belokobylskij, 1992

259(243).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... Aivalykus Nixon, 1938
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... Leptospathius Szépligeti, 1902
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long... Australospathius Belokobylskij, Iqbal & Austin, 2004

260(242).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 261
  • Brachial (1st subdiscal) cell of fore wing open... 278

261(260).
  • First and second tergites not fused ... 262
  • First and second tergites fused... 277

262(261).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... 263
  • Hind femora not swollen (at least 3x longer than wide)... 264

263(262).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeum almost entirely smooth (visible in the carinated areola); ovipositor nodus double, with a second node weakly developed; mesoscutum surface of median and lateral lobes smooth and shining ... Ceylonspathius Belokobylskij, 2002
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; propodeum fully or partly (micro)sculptured; ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); mesoscutum surface of median and lateral lobes more or less sculptured... Platyspathius Viereck, 1911

264(262).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... 265
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A)... 269
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 271

265(264).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 266
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)

266(265).
  • First flagellar segment longer than second ... 267
  • First flagellar segment equal to or shorter than second... 268

267(266).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... Caenophanes Foerster, 1862 (including the subgenus)
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... Chelonodoryctes Belokobylskij & Quicke, 2000
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long... Spathius Nees, 1918

268(266).
  • Occipital carina meeting hypostomal carina; propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); fore tibial spines more or less numerous and dispersed; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area) ... Rhacontsira Belokobylskij, 1998
  • Occipital carina not meeting hypostomal carina; propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); fore tibial spines usually few and forming a single row; third tergite medially without any distinct transverse narrow depression/furrow... Eodendrus Belokobylskij, 1998

269(264).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; dorsope of first tergite present and more or less distinct ... 270
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

270(269).
  • Sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body); fore wing of male with sclerotized enlargement; fore femur without anterodorsal protuberances ... Neurocrassus Snoflak, 1945
  • Sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); metapleuron glabrous or with a very sparse/rare discret pubescence; fore wing of male without sclerotized enlargement, including veins 1-m and 1-SR+m; fore femur with more or less distinct anterodorsal protuberances... Ipodoryctes Granger, 1949

271(264).
  • First tergite without basolateral process (pointed wing-like, spine-like projections) near base ... 272
  • First tergite with basolateral process (pointed wing-like, spine-like projections) near base... 276

272(271).
  • First tergite without dorsal carinae ... Halycaea Cameron, 1903
  • First tergite with dorsal carina (one or two, often mediolongitudinally or submediolongitudinally)(if the tergite is fully longitudinally striate, this(these) carina(e) could be hardly distinguished from the other and are coded as present)... 273

273(272).
  • Second radiomedial vein (r-m) of fore wing with wide bulla ... 274
  • Second radiomedial vein (r-m) of fore wing largely tubular... Pedinotus Szépligeti, 1902 (including the subgenus Eopedinotus)

274(273).
  • First tergite from 1 times to 1.5 times longer than apical width ... 275
  • First tergite from 1.5 times to 2.5 times as long its apical width... Hypodoryctes Kokujev, 1900

275(274).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances ... Doryctes Haliday, 1836
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances... Ontsira Cameron, 1900

276(271).
  • First flagellar segment longer than second; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; occipital carina meeting hypostomal carina; propodeum with lateral long spines (higher than its base); first tergite from 1.5 times to 2.5 times as long its apical width ... Ryukyuspathius Belokobylskij, 2008
  • First flagellar segment equal to or shorter than second; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; occipital carina not meeting hypostomal carina; propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width... Rasnitsynoryctes Belokobylskij, 2011

277(261).
  • First flagellar segment longer than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); fore tibial spines more or less numerous and dispersed; third tergite medially without any distinct transverse narrow depression/furrow; mid femora without anterodorsal protuberances ... Mimipodoryctes Belokobylskij, 2000
  • First flagellar segment equal to or shorter than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines usually few and forming a single row; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); mid femora with more or less distinct anterodorsal protuberances... Arhaconotus Belokobylskij, 2000

278(260).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 279
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... Hecabalodes Wilkinson, 1929

279(278).
  • Occipital carina meeting hypostomal carina ... 280
  • Occipital carina not meeting hypostomal carina... 283

280(279).
  • Second radiomedial vein (r-m) of fore wing present ... 281
  • Second radiomedial vein (r-m) of fore wing absent... Monolexis Foerster, 1862

281(280).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scape shorter than first flagellomere; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; propodeum convex or abruptly declivous posteriorly ... Heterospilus Halliday, 1836 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); scape equal to or longer than first flagellomere; first tergite apical width more than 1.6 times (even 2 times) basal width; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... 282

282(281).
  • First flagellar segment longer than second; propodeum without lateral tubercles; eyes glabrous; pronotum with convex lobe; scutellum not flattened (e.g. swollen, conical, sharply raised) ... Neoheterospilus Belokobylskij, 2006 (including the subgenera)
  • First flagellar segment equal to or shorter than second; propodeum with lateral tubercles (as high as large at base); eyes setose; pronotum dorsally without modifications; scutellum flat... Spathiomorpha Tobias, 1976

283(279).
  • Ovipositor nodus without visible node (but lower valves serrate) ... Hecabolomorpha Belokobylskij & Chen, 2006
  • Ovipositor nodus double, with a second node weakly developed... Ceylonspathius Belokobylskij, 2002
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak)... 284

284(283).
  • Propodeal carinated areas present at least basally; hind femora not swollen (at least 3x longer than wide); mesoscutum surface of median and lateral lobes smooth and shining; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); hind wing male without a stigma-like enlargement ... Parallorhogas Marsh, 1993
  • Propodeal carinated areas completely absent; hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); mesoscutum surface of median and lateral lobes more or less sculptured; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts... Hemidoryctes Belokobylskij, 1993

285(241).
  • Propodeal bridge between abdominal and coxal foramina absent; occipital carina present, at least laterally; malar suture present; pronotum dorsally without modifications; epicnemial carina present ... 286
  • Propodeal bridge between abdominal and coxal foramina present; occipital carina absent; malar suture absent; pronotum with convex lobe; epicnemial carina absent... Siragra Cameron, 1907

286(285).
  • Brachial (1st subdiscal) cell of fore wing distally closed; third tergite medially without any distinct transverse narrow depression/furrow; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; frons not excavated (flat or convex) ... 287
  • Brachial (1st subdiscal) cell of fore wing open; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); hind basitarsus up to 0.45 times as long as tarsi 2-5 combined; frons excavated (concave, sometimes weakly)... Synspilus Belokobylskij & Quicke, 2000

287(286).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); first tergite from 1.5 times to 2.5 times as long its apical width ... Arkoola Belokobylskij, Iqbal & Austin, 2004
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); first tergite more 2.5 times its apical width... Paraspathius Nixon, 1943

288(1).
  • Apical lobe of scape absent ... 289
  • Apical lobe of scape present... 331

289(288).
  • Basoventral tooth of hind coxa absent ... 290
  • Basoventral tooth of hind coxa present (sometimes very small)... 306

290(289).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum ... 291
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum... 301

291(290).
  • Frons without lateral protuberances protruding to the surface of head ... 292
  • Frons with lateral protuberances protruding to the surface of head... 300

292(291).
  • Fore tibial spines more or less numerous and dispersed ... 293
  • Fore tibial spines usually few and forming a single row... 294

293(292).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; first tergite from 1 times to 1.5 times longer than apical width; mesoscutum surface of median and lateral lobes more or less sculptured; ocelar triangle with base shorter than its sides; labial palpi one-segmented ... Nipponecphylus Belokobylskij & Konishi, 2001 (females)
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; first tergite from 1.5 times to 2.5 times as long its apical width; mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa); ocelar triangle equilateral; labial palpi three-segmented... Fijibracon Belokobylskij, 1995
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; first tergite more 2.5 times its apical width; mesoscutum surface of median and lateral lobes smooth and shining; ocelar triangle with base larger than its sides; labial palpi four-segmented... Fijispathius Belokobylskij, Iqbal & Austin, 2004

294(292).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; third labial palp segment as long as or longer than second ... 295
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; third labial palp segment distinctly shorter than second... 299

295(294).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... 296
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A)... Sonanus Belokobylskij & Konishi, 2001
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 298

296(295).
  • First flagellar segment longer than second; occipital carina meeting hypostomal carina; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); hind tibia without comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves absent ... 297
  • First flagellar segment equal to or shorter than second; occipital carina not meeting hypostomal carina; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); hind tibia with comb of dense flat setae at inner apex; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... Eodendrus Belokobylskij, 1998

297(296).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; propodeal carinated areas completely absent; propodeum without lateral tubercles; first tergite from 1 times to 1.5 times longer than apical width; first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view ... Euscelinus Westwood, 1882
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; propodeal carinated areas present at least basally; propodeum with lateral tubercles (as high as large at base); first tergite from 1.5 times to 2.5 times as long its apical width; first flagellar segment surface regularly smooth or finely sculptured... Spathius (Ambispathius) Belokobylskij, 1995

298(295).
  • First flagellar segment longer than second; propodeal carinated areas completely absent; propodeum without lateral tubercles; hind femora not swollen (at least 3x longer than wide); first tergite from 1 times to 1.5 times longer than apical width ... Bracodoryctes Belokobylskij & Quicke, 2000
  • First flagellar segment equal to or shorter than second; propodeal carinated areas present at least basally; propodeum with lateral tubercles (as high as large at base); hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); first tergite less than 1 times longer than apical width... Cryptontsira Belokobylskij, 2008

299(294).
  • First tergite less than 1 times longer than apical width; malar suture present; notauli entirely absent; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); scape shorter than first flagellomere ... Liodoryctes Szépligeti, 1906
  • First tergite from 1 times to 1.5 times longer than apical width; malar suture absent; notauli complete or at least partly present (smooth or sculptured); sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); scape equal to or longer than first flagellomere... Zombrus Marshall, 1897

300(291).
  • First flagellar segment longer than second; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); pronotum with pointed spines or tubercles; hind wing male without a stigma-like enlargement ... Ceratodoryctes Belokobylskij, Iqbal & Austin, 2004
  • First flagellar segment equal to or shorter than second; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; pronotum dorsally without modifications; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts... Dendrosoter Wesmael, 1838 (including the subgenus)

301(290).
  • Hind wing with the recurrent vein (m-cu); fourth-sixth metasomal tergites of males smooth (or striate/coriaceous) basaly ... 302
  • Hind wing without the recurrent vein (m-cu); fourth-sixth metasomal tergites of males at least crenulate-striate basally (often on all their surface)... 305

302(301).
  • Recurrent vein (m-cu) of hind wing not curved towards apex of wing; propodeal bridge between abdominal and coxal foramina absent; first flagellar segment longer than second; hind wing without transverse vein (r) of radial cell; fore femur without anterodorsal protuberances ... 303
  • Recurrent vein (m-cu) of hind wing clearly (sometimes strongly) curved to apex of wing; propodeal bridge between abdominal and coxal foramina present; first flagellar segment equal to or shorter than second; hind wing with a transverse vein (r) of radial cell; fore femur with more or less distinct anterodorsal protuberances... Leptospathius Szépligeti, 1902

303(302).
  • Propodeal carinated areas present at least basally; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); first flagellar segment surface regularly smooth or finely sculptured; hind wing male without a stigma-like enlargement; dorsal spines of hind tibia absent ... 304
  • Propodeal carinated areas completely absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; dorsal spines of hind tibia present... Doryctophasmus Enderlein, 1912

304(303).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; eyes setose; dorsope of first tergite present and more or less distinct ... Rhoptrocentrus Marshall, 1897
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; eyes glabrous; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

305(301).
  • First flagellar segment longer than second; second radiomedial vein (r-m) of fore wing present; brachial (1st subdiscal) cell of fore wing distally closed; hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak) ... Acanthodoryctes Turner, 1918
  • First flagellar segment equal to or shorter than second; second radiomedial vein (r-m) of fore wing absent; brachial (1st subdiscal) cell of fore wing open; hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); ovipositor nodus double, with a second node weakly developed... Spathiostenus Belokobylskij, 1992

306(289).
  • Frons without lateral protuberances protruding to the surface of head ... 307
  • Frons with lateral protuberances protruding to the surface of head... Ceratodoryctes Belokobylskij, Iqbal & Austin, 2004

307(306).
  • First and second tergites not fused ... 308
  • First and second tergites fused... 330

308(307).
  • Wings present ... 309
  • Wings absent... 329

309(308).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 310
  • Brachial (1st subdiscal) cell of fore wing open... 323

310(309).
  • Third tergite medially without any distinct transverse narrow depression/furrow ... 311
  • Third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area)... 321

311(310).
  • Propodeum fully or partly (micro)sculptured; first flagellar segment surface regularly smooth or finely sculptured ... 312
  • Propodeum almost entirely smooth (visible in the carinated areola); first flagellar segment surface smooth dorsally, more or less sculptured ventrally, strongly compressed dorso-ventrally and curved in lateral view, often ovale in frontal view... Dooliba Belokobylskij, Iqbal & Austin, 2004

312(311).
  • Hind wing without transverse vein (r) of radial cell; scape less than two times longer than its maximum width ... 313
  • Hind wing with a transverse vein (r) of radial cell; scape two or more times longer than its maximum width... Pirramurra Belokobylskij, Iqbal & Austin, 2004

313(312).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Platyspathius Viereck, 1911
  • Hind femora not swollen (at least 3x longer than wide)... 314

314(313).
  • Occipital carina present, at least laterally; epicnemial carina present; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded) ... 315
  • Occipital carina absent; epicnemial carina absent; propodeum convex or abruptly declivous posteriorly... Bracodoryctes Belokobylskij & Quicke, 2000

315(314).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial ... 316
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A)... 318
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A)... 319

316(315).
  • First flagellar segment longer than second; mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; propodeal carinated areas present at least basally; basal area of second tergite enclosed by carinae or grooves absent ... 317
  • First flagellar segment equal to or shorter than second; mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; propodeal carinated areas completely absent; basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves)... Eodendrus Belokobylskij, 1998

317(316).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; dorsope of first tergite present and more or less distinct ... Caenophanes Foerster, 1862 (including the subgenus)
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; first tergite apical width more than 1.6 times (even 2 times) basal width; dorsope of first tergite absent or indistinct... Spathius Nees, 1918

318(315).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present; scutellum not flattened (e.g. swollen, conical, sharply raised); basal area of second tergite enclosed by carinae or grooves present (sometimes shallow grooves) ... Ontsirospathius Belokobylskij, Iqbal & Austin, 2004
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; scutellum flat; basal area of second tergite enclosed by carinae or grooves absent... Spathius Nees, 1918

319(315).
  • Metapleuron glabrous or with a very sparse/rare discret pubescence; first tergite with dorsal carina (one or two, often mediolongitudinally or submediolongitudinally)(if the tergite is fully longitudinally striate, this(these) carina(e) could be hardly distinguished from the other and are coded as present) ... 320
  • Metapleuron with clear long setae (sparse or numerous), sometimes dense pubescence (in this case often present on other part of body); first tergite without dorsal carinae... Halycaea Cameron, 1903

320(319).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances ... Doryctes Haliday, 1836
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances... Ontsira Cameron, 1900

321(310).
  • Pronotum with convex lobe; mesoscutum surface of median lobe sculptured and lateral lobes smooth (or vice versa); fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances; hind femora without anterodorsal protuberances ... Neorhaconotus Belokobylskij, Iqbal & Austin, 2004
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances; hind femora with more or less distinct anterodorsal protuberances... 322

322(321).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines more or less numerous and dispersed; sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); scape shorter than first flagellomere; hind tibia without comb of dense flat setae at inner apex ... Rhacontsira Belokobylskij, 1998
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); fore tibial spines usually few and forming a single row; sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); scape equal to or longer than first flagellomere; hind tibia with comb of dense flat setae at inner apex... Ipodoryctes Granger, 1949

323(309).
  • Occipital carina meeting hypostomal carina ... 324
  • Occipital carina not meeting hypostomal carina... 327

324(323).
  • Second radiomedial vein (r-m) of fore wing present ... 325
  • Second radiomedial vein (r-m) of fore wing absent... Monolexis Foerster, 1862

325(324).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); scape shorter than first flagellomere; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; propodeum convex or abruptly declivous posteriorly ... Heterospilus Halliday, 1836 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); scape equal to or longer than first flagellomere; first tergite apical width more than 1.6 times (even 2 times) basal width; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... 326

326(325).
  • First flagellar segment longer than second; propodeum without lateral tubercles; pronotum with convex lobe; scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron) ... Neoheterospilus Belokobylskij, 2006 (including the subgenera)
  • First flagellar segment equal to or shorter than second; propodeum with lateral tubercles (as high as large at base); pronotum dorsally without modifications; scutellum flat; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron)... Spathiomorpha Tobias, 1976

327(323).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs) ... Hemidoryctes Belokobylskij, 1993
  • Hind femora not swollen (at least 3x longer than wide)... 328

328(327).
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes more or less sculptured; hind wing male with a stigma-like enlargement in distal part of costal (1-SC+R) vein, without incurved marginal parts; fore femur without anterodorsal protuberances; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Leluthia Cameron, 1887 (including the subgenus)
  • Parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); mesoscutum surface of median and lateral lobes smooth and shining; hind wing male without a stigma-like enlargement; fore femur with more or less distinct anterodorsal protuberances; first tergite apical width more than 1.6 times (even 2 times) basal width... Parallorhogas Marsh, 1993

329(308).
  • First flagellar segment longer than second; propodeum almost entirely smooth (visible in the carinated areola); propodeum with lateral long spines (higher than its base); eyes setose; notauli complete or at least partly present (smooth or sculptured) ... Echinodoryctes Belokobylskij, Iqbal & Austin, 2004
  • First flagellar segment equal to or shorter than second; propodeum fully or partly (micro)sculptured; propodeum with lateral tubercles (as high as large at base); eyes glabrous; notauli entirely absent... Doryctopsis Belokobylskij, Iqbal & Austin, 2004

330(307).
  • First flagellar segment longer than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); fore tibial spines more or less numerous and dispersed; third tergite medially without any distinct transverse narrow depression/furrow; mid femora without anterodorsal protuberances ... Mimipodoryctes Belokobylskij, 2000
  • First flagellar segment equal to or shorter than second; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; fore tibial spines usually few and forming a single row; third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); mid femora with more or less distinct anterodorsal protuberances... Arhaconotus Belokobylskij, 2000

331(288).
  • Frons without lateral protuberances protruding to the surface of head; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... 332
  • Frons with lateral protuberances protruding to the surface of head; first tergite apical width more than 1.6 times (even 2 times) basal width... Ceratodoryctes Belokobylskij, Iqbal & Austin, 2004

332(331).
  • Propodeal bridge between abdominal and coxal foramina absent; epicnemial carina present; frons not excavated (flat or convex) ... 333
  • Propodeal bridge between abdominal and coxal foramina present; epicnemial carina absent; frons excavated (concave, sometimes weakly)... Siragra Cameron, 1907

333(332).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent; recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); hind tibia without comb of dense flat setae at inner apex; first tergite with dorsal carina (one or two, often mediolongitudinally or submediolongitudinally)(if the tergite is fully longitudinally striate, this(these) carina(e) could be hardly distinguished from the other and are coded as present) ... 334
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long; recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); hind tibia with comb of dense flat setae at inner apex; first tergite without dorsal carinae... Paraspathius Nixon, 1943

334(333).
  • Basoventral tooth of hind coxa absent; propodeal carinated areas completely absent ... 335
  • Basoventral tooth of hind coxa present (sometimes very small); propodeal carinated areas present at least basally... 336

335(334).
  • Hind wing with a small or medium sized submedial (subbasal) cell, the first abscissa of M+CU 0.2-1.0 times as long as second abscissa (1-M); third tergite with a distinct transverse narrow furrow usually between its anterior third and the middle (sometimes meeting laterally second metasomal suture and enclosing a lenticular area); pronotum dorsally without modifications; sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated) ... Syngaster Brullé, 1846
  • Hind wing with a distinctly enlarged submedial (subbasal) cell, the first abscissa of M+CU longer, often 1.5-2.0 times, than the second abscissa (1-M); third tergite medially without any distinct transverse narrow depression/furrow; pronotum with pointed spines or tubercles; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully)... Antidoryctes Belokobylskij & Quicke, 2000

336(334).
  • Propodeum without lateral tubercles; dorsal spines of hind tibia present; hind basitarsus 0.75 times as long as tarsi 2-5 combined or more; first tergite with basolateral process (pointed wing-like, spine-like projections) near base; first tergite median carinae very short (less than or equal to the third of petiole length) ... 337
  • Propodeum with lateral tubercles (as high as large at base); dorsal spines of hind tibia absent; hind basitarsus 0.45 to 0.75 times as long as tarsi 2-5 combined; first tergite without basolateral process (pointed wing-like, spine-like projections) near base; first tergite median carinae reaching, or nearly, the half length of the petiole... Jarra Marsh & Austin, 1994

337(336).
  • Nervellus (cu-a) of fore wing postfurcal or interstitial; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); sternaulus (precoxal sulcus) without sculptures (may be smooth or coriaceous, but not carinated); second radiomedial vein (r-m) of fore wing with wide bulla; subapical teeth on fore and middle femora ventrally present ... Pseudosyngaster Belokobylskij, Iqbal & Austin, 2004 (female)
  • Nervellus (cu-a) of fore wing antefurcal; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising before or from middle of distal vein of brachial cell (large space between the curved part and the level of the vein 2-1A); sternaulus (precoxal sulcus) at least partly sculptured (even only anteriorly; sometimes fully); second radiomedial vein (r-m) of fore wing largely tubular; subapical teeth on fore and middle femora ventrally absent... Gurburra Belokobylskij, Iqbal & Austin, 2004

338(1).
  • Basoventral tooth of hind coxa absent ... 339
  • Basoventral tooth of hind coxa present (sometimes very small)... 344

339(338).
  • Acrosternite of first tergite short (shorter than 0.25 as long as the tergite, fused or not with ventral margins of tergite), petiole absent ... 340
  • Acrosternite of first tergite long (0.3-0.5 as long as the tergite, fused with its ventral margin anteriorly), petiole present... 341
  • Acrosternite of first tergite very long (more than 0.52 times as long as tergite, sometimes entirely fused with it ventral margin) petiole present and long... 343

340(339).
  • First flagellar segment longer than second; propodeal carinated areas completely absent; propodeum without lateral tubercles; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; first tergite from 1 times to 1.5 times longer than apical width ... Euscelinus Westwood, 1882
  • First flagellar segment equal to or shorter than second; propodeal carinated areas present at least basally; propodeum with lateral tubercles (as high as large at base); parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); first tergite less than 1 times longer than apical width... Cryptontsira Belokobylskij, 2008

341(339).
  • First flagellar segment longer than second; occipital carina absent; scape two or more times longer than its maximum width; labial palpi four-segmented ... Sisupala Nixon, 1943
  • First flagellar segment equal to or shorter than second; occipital carina present, at least laterally; scape less than two times longer than its maximum width; labial palpi three-segmented... 342

342(341).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; propodeal carinated areas present at least basally; fore tibial spines more or less numerous and dispersed; first tergite from 1.5 times to 2.5 times as long its apical width; scutellum flat ... Fijibracon Belokobylskij, 1995
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; propodeal carinated areas completely absent; fore tibial spines usually few and forming a single row; first tergite from 1 times to 1.5 times longer than apical width; scutellum not flattened (e.g. swollen, conical, sharply raised)... Ecphylopsis Ashmead, 1900

343(339).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; propodeum with lateral long spines (higher than its base); propodeum almost entirely smooth (visible in the carinated areola); first tergite apical width less than or equal to 1.5 times (or subequal to) basal width; eyes setose ... Fijispathius Belokobylskij, Iqbal & Austin, 2004
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; propodeum without lateral tubercles; propodeum fully or partly (micro)sculptured; first tergite apical width more than 1.6 times (even 2 times) basal width; eyes glabrous... Spathius Nees, 1918

344(338).
  • Second radiomedial vein (r-m) of fore wing present ... 345
  • Second radiomedial vein (r-m) of fore wing absent... 352

345(344).
  • Brachial (1st subdiscal) cell of fore wing distally closed ... 346
  • Brachial (1st subdiscal) cell of fore wing open... 349

346(345).
  • Fifth or sixth metasomal tergites not enlarged, nor covering succeeding tergites, entirely smooth ... 347
  • Fifth or sixth metasomal tergites more or less distinctly enlarged, covering succeeding tergites and sculptured at least basally... Rhaconotus Ruthe, 1854 (including the subgenus Pararhacon)

347(346).
  • Recurrent vein (m-cu) of fore wing postfurcal (arising distad 2RS, (RS+M)b absent); dorsope of first tergite absent or indistinct ... Spathius Nees, 1918
  • Recurrent vein (m-cu) of fore wing interstitial or antefurcal (arising basad or directly in line with 2RS, (RS+M)b present then second submarginal cell petiolate horizontally); dorsope of first tergite present and more or less distinct... 348

348(347).
  • Pronotum with convex lobe; mesoscutum surface of median and lateral lobes smooth and shining; fore femur without anterodorsal protuberances; mid femora without anterodorsal protuberances ... Doryctes Haliday, 1836
  • Pronotum dorsally without modifications; mesoscutum surface of median and lateral lobes more or less sculptured; fore femur with more or less distinct anterodorsal protuberances; mid femora with more or less distinct anterodorsal protuberances... Ontsira Cameron, 1900

349(345).
  • Hind femora swollen, at most 2.5x longer than wide (often also the case for fore and mid legs); notauli entirely absent; scape equal to or longer than first flagellomere; propodeum convex or abruptly declivous posteriorly ... Stenocorse Marsh, 1968
  • Hind femora not swollen (at least 3x longer than wide); notauli complete or at least partly present (smooth or sculptured); scape shorter than first flagellomere; propodeum not convex, sloping from base to apex or horizontal, to weakly convex (only slowly rounded)... 350

350(349).
  • Ovipositor nodus double, with a second node weakly developed; fore femur without anterodorsal protuberances; first tergite apical width less than or equal to 1.5 times (or subequal to) basal width ... Glyptocolastes Ashmead, 1900
  • Ovipositor nodus double (or more), with both nodus developed at the same level than the other (even if weak); fore femur with more or less distinct anterodorsal protuberances; first tergite apical width more than 1.6 times (even 2 times) basal width... 351

351(350).
  • Occipital carina meeting hypostomal carina; eyes setose; mesoscutum surface of median and lateral lobes more or less sculptured; scutellum not flattened (e.g. swollen, conical, sharply raised); sternaulus (precoxal sulcus) long (more than 2/3 length of mesopleuron) ... Doryctinus Roman 1910
  • Occipital carina not meeting hypostomal carina; eyes glabrous; mesoscutum surface of median and lateral lobes smooth and shining; scutellum flat; sternaulus (precoxal sulcus) shortened (less or, at most, equal to 2/3 length of mesopleuron)... Parallorhogas Marsh, 1993

352(344).
  • Mesoscutum with a sharply raised face anteriorly, at right angle with pronotum; brachial (1st subdiscal) cell of fore wing open; occipital carina meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing not interstitial, arising behind middle of distal vein of brachial cell (short space between the curved part and the level of the vein 2-1A); scutellum flat ... Monolexis Foerster, 1862
  • Mesoscutum without a sharply raised face anteriorly, sometimes on nearly same plane as pronotum; brachial (1st subdiscal) cell of fore wing distally closed; occipital carina not meeting hypostomal carina; parallel vein (CU1a) relative to cubital vein (2-CU1) of fore wing interstitial; scutellum not flattened (e.g. swollen, conical, sharply raised)... Kauriphanes Belokobylskij & Zaldívar, 2011

Cite this publication as: ‘L. Watson and M. J. Dallwitz (1998 onwards). DELTA Sample Data: Descriptions, Illustrations, Identification, and Information Retrieval. Version: 21st September 2000. http://biodiversity.uno.edu/delta/’. Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993, 1995, 2000) should also be cited (see References).

Index

Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith